Ctenochasmatidae

Reconstruction of Gnathosaurus subulatus, copyright Dmitry Bogdanov.

Belongs within: Pterodactyloidea.

The Ctenochasmatidae are a group of pterosaurs known from the Upper Jurassic to the mid-Cretaceous, characterised by elongate jaws bearing high numbers of slender teeth (Witton 2013).

<==Ctenochasmatidae [Ptenodraconinae, Pterodaustridae, Pterodaustriidae]
    |  i. s.: Gegepterus changiRK13, W13
    |         Beipiaopterus Lü 2003A08
    |           `--B. chenianusW13
    |--Ctenochasmatinae [Ctenochasminae]U03
    |    |--Eosipterus Ji & Ji 1997U03
    |    |    `--E. yangiW13
    |    |--Pterodaustro Bonaparte 1970U03 [incl. Puntanipterus Bonaparte & Sanchez 1975W91]
    |    |    `--P. guinazui Bonaparte 1970K03 [incl. Puntanipterus globosus Bonaparte & Sanchez 1975W13, U03]
    |    `--Ctenochasma von Meyer 1852A08
    |         |--C. elegans (Wagner 1861)W13, A08 [=Pterodactylus elegansU03; incl. C. porocristata Buisonjé 1981W13, W91]
    |         |--C. gracile Oppel 1862K03
    |         |--C. roemeri Meyer 1851W91
    |         `--C. taquetiW13
    `--GnathosaurinaeU03
         |--Ardeadactylus longicollum (Meyer 1854)RK13, U03 [=Pterodactylus longicollumU03; incl. Auroazhdarcho primordiusW13]
         |--Diopecephalus Seeley 1871U03
         |--Huanhepterus Dong 1982U03
         |    `--H. quingyangensis Dong 1982B93
         |--Plataleorhynchus Howse & Milner 1995U03
         |    `--P. streptorophodonW13
         `--Gnathosaurus von Meyer 1834 [incl. Doratorhynchus Seeley 1875, Ptenodracon Lydekker 1888]A08
              |--G. macrurusW13 [=Pterodactylus macrurusRK13]
              |--‘Pterodactylus’ micronyx von Meyer 1856A08
              |--G. subulatus Meyer 1834W91
              `--‘Doratorhynchus’ validus Seeley 1875PSB03 (see below for synonymy)

‘Doratorhynchus’ validus Seeley 1875PSB03 [=Doratohynchus validum (l. c.)RK13, Ornithocheirus validusRK13, Pterodactylus validus Owen 1874 (n. n.)RK13]

*Type species of generic name indicated

References

[A08] Averianov, A. O. 2008. Superorder Pterosauria. In: Ivakhnenko, M. F., & E. N. Kurochkin (eds) Fossil Vertebrates of Russia and Adjacent Territories. Fossil Reptiles and Birds. Part 1 pp. 319–342. GEOS: Moscow.

[B93] Benton, M. J. 1993. Reptilia. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 681–715. Chapman & Hall: London.

[K03] Kellner, A. W. A. 2003. Pterosaur phylogeny and comments on the evolutionary history of the group. In: Buffetaut, E., & J.-M. Mazin (eds) Evolution and Palaeobiology of Pterosaurs. Geological Society Special Publications 217: 105–137. The Geological Society: London.

[PSB03] Pereda Suberbiola, X., N. Bardet, S. Jouve, M. Iarochène, B. Bouya & M. Amaghzaz. 2003. A new azhdarchid pterosaur from the Late cretaceous phosphates of Morocco. In: Buffetaut, E., & J.-M. Mazin (eds) Evolution and Palaeobiology of Pterosaurs. Geological Society Special Publications 217: 79–90. The Geological Society: London.

[RK13] Rodrigues, T., & A. W. A. Kellner. 2013. Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England. ZooKeys 308: 1–112.

[U03] Unwin, D. M. 2003. On the phylogeny and evolutionary history of pterosaurs. In: Buffetaut, E., & J.-M. Mazin (eds) Evolution and Palaeobiology of Pterosaurs. Geological Society Special Publications 217: 139–190. The Geological Society: London.

[W91] Wellnhofer, P. 1991. The Illustrated Encyclopedia of Pterosaurs. Salamander Books: London (reprinted in: Norman, D., & P. Wellnhofer. 2000. The Illustrated Encyclopedia of Dinosaurs. Salamander Books).

[W13] Witton, M. P. 2013. Pterosaurs: Natural History, Evolution, Anatomy. Princeton University Press: Princeton (New Jersey).

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