
Belongs within: Pterodactyloidea.
The Ctenochasmatidae are a group of pterosaurs known from the Upper Jurassic to the mid-Cretaceous, characterised by elongate jaws bearing high numbers of slender teeth (Witton 2013).
<==Ctenochasmatidae [Ptenodraconinae, Pterodaustridae, Pterodaustriidae] | i. s.: Gegepterus changiRK13, W13 | Beipiaopterus Lü 2003A08 | `--B. chenianusW13 |--Ctenochasmatinae [Ctenochasminae]U03 | |--Eosipterus Ji & Ji 1997U03 | | `--E. yangiW13 | |--Pterodaustro Bonaparte 1970U03 [incl. Puntanipterus Bonaparte & Sanchez 1975W91] | | `--P. guinazui Bonaparte 1970K03 [incl. Puntanipterus globosus Bonaparte & Sanchez 1975W13, U03] | `--Ctenochasma von Meyer 1852A08 | |--C. elegans (Wagner 1861)W13, A08 [=Pterodactylus elegansU03; incl. C. porocristata Buisonjé 1981W13, W91] | |--C. gracile Oppel 1862K03 | |--C. roemeri Meyer 1851W91 | `--C. taquetiW13 `--GnathosaurinaeU03 |--Ardeadactylus longicollum (Meyer 1854)RK13, U03 [=Pterodactylus longicollumU03; incl. Auroazhdarcho primordiusW13] |--Diopecephalus Seeley 1871U03 |--Huanhepterus Dong 1982U03 | `--H. quingyangensis Dong 1982B93 |--Plataleorhynchus Howse & Milner 1995U03 | `--P. streptorophodonW13 `--Gnathosaurus von Meyer 1834 [incl. Doratorhynchus Seeley 1875, Ptenodracon Lydekker 1888]A08 |--G. macrurusW13 [=Pterodactylus macrurusRK13] |--‘Pterodactylus’ micronyx von Meyer 1856A08 |--G. subulatus Meyer 1834W91 `--‘Doratorhynchus’ validus Seeley 1875PSB03 (see below for synonymy)
‘Doratorhynchus’ validus Seeley 1875PSB03 [=Doratohynchus validum (l. c.)RK13, Ornithocheirus validusRK13, Pterodactylus validus Owen 1874 (n. n.)RK13]
*Type species of generic name indicated
References
[A08] Averianov, A. O. 2008. Superorder Pterosauria. In: Ivakhnenko, M. F., & E. N. Kurochkin (eds) Fossil Vertebrates of Russia and Adjacent Territories. Fossil Reptiles and Birds. Part 1 pp. 319–342. GEOS: Moscow.
[B93] Benton, M. J. 1993. Reptilia. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 681–715. Chapman & Hall: London.
[K03] Kellner, A. W. A. 2003. Pterosaur phylogeny and comments on the evolutionary history of the group. In: Buffetaut, E., & J.-M. Mazin (eds) Evolution and Palaeobiology of Pterosaurs. Geological Society Special Publications 217: 105–137. The Geological Society: London.
[PSB03] Pereda Suberbiola, X., N. Bardet, S. Jouve, M. Iarochène, B. Bouya & M. Amaghzaz. 2003. A new azhdarchid pterosaur from the Late cretaceous phosphates of Morocco. In: Buffetaut, E., & J.-M. Mazin (eds) Evolution and Palaeobiology of Pterosaurs. Geological Society Special Publications 217: 79–90. The Geological Society: London.
[RK13] Rodrigues, T., & A. W. A. Kellner. 2013. Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England. ZooKeys 308: 1–112.
[U03] Unwin, D. M. 2003. On the phylogeny and evolutionary history of pterosaurs. In: Buffetaut, E., & J.-M. Mazin (eds) Evolution and Palaeobiology of Pterosaurs. Geological Society Special Publications 217: 139–190. The Geological Society: London.
[W91] Wellnhofer, P. 1991. The Illustrated Encyclopedia of Pterosaurs. Salamander Books: London (reprinted in: Norman, D., & P. Wellnhofer. 2000. The Illustrated Encyclopedia of Dinosaurs. Salamander Books).
[W13] Witton, M. P. 2013. Pterosaurs: Natural History, Evolution, Anatomy. Princeton University Press: Princeton (New Jersey).