Cyprois marginata, copyright Zoologische Staatssammlung Muenchen.

Belongs within: Ostracoda.
Contains: Suchonellina, Cyprididae, Candonidae, Ilyocyprididae, Cyprideidae.

The Cypridocopina are a group of ostracods in which three pairs of thoracic legs are present, with the first pair modified to function as a mouthpart and the third pair is bent dorsally and used as a cleaning organ, and the body bears a terminal furca (Benson et al. 1961). The majority of species have a smooth carapace though exceptions occur (Liebau 2005). Members of the superfamily Macrocypridoidea have a central muscle scar pattern of about nine scars forming a loosely aggregated group with a further three scars more dorsad, whereas Pontocyprididae have a ‘paw-print’ pattern of three scars in front of a fourth.

Characters (from Benson et al. 1961, as Cypridacea): Carapace variable in size, shape, and surface ornamentation; calcareous or corneous; mid-ventral margin curved inward. Hinge of ridge-and-groove type or rabbeted; muscle scars a median group of spots, typically not radially or linearly arranged, with additional more anteroventral spots in most genera; inner lamellae present; line of concrescence and inner margin typically separated.

    |--Pontocyprididae [Argilloeciinae, Clinocyprinae, Pontocypridae, Pontocypridinae, Pontocypridoidea, Pontocyprinae]MD01
    |    |--Liasina lanceolata (Apostolecsu 1959)WSB93
    |    |--Argilloecia Sars 1866 [=Argillaecia Brady 1870]BB61
    |    |    `--*A. cylindrica Sars 1866BB61
    |    |--Bakunella Schneider in Mandelstam et al. 1958BB61
    |    |    |--*B. dorsoarcuata (Zalanyi 1929) [=Pontocypris dorsoarcuata]BB61
    |    |    `--B. abkhazica Vekua 1965CC02
    |    |--Clinocypris Mandelstam 1956BB61
    |    |    `--*C. scolia Mandelstam 1956BB61
    |    |--Pontocypria Müller 1894BB61
    |    |    `--*P. spinosa Müller 1894BB61
    |    |--Propontocypris Sylvester-Bradley 1947BB61
    |    |    `--*P. trigonella (Sars 1866) [=Pontocypris trigonella]BB61
    |    |--Subulacypris Schneider in Mandelstam et al. 1957BB61
    |    |    `--*S. subtilis Schneider in Mandelstam et al. 1957BB61
    |    |--Bosquetia Brady, Crosskey & Robertson 1874M61, BB61
    |    |    `--*B. robusta Brady, Crosskey & Robertson 1874BB61
    |    `--Pontocypris Sars 1866 [incl. Cytheropsis McCoy 1849 (n. d.), Erythrocypris Müller 1894]BB61
    |         |--P. mytiloides (Norman 1862) (see below for synonymy)BB61
    |         |--*Cytheropsis’ aldensis M’Coy 1851BB61
    |         `--P. mawii Jones 1887 [incl. P. mawii var. proxima Jones 1889]C01
    |    |--Macrocyprididae [Macrocypridae, Macrocyprinae]WSB93
    |    |    |--Macrosarisa exquisita (Kaye 1964)WSB93
    |    |    |--Macrocypris Brady 1867CB86
    |    |    |    |--*M. minna (Baird 1850) [=Cythere minna]BB61
    |    |    |    `--M. siliquoides Jones 1887C01
    |    |    `--Macrocypria Sars 1923BB61
    |    |         `--*M. sarsi (Müller 1912) (see below for synonymy)BB61
    |    `--AcratiidaeL05
    |         |--Acratina Egorov 1953L05, BB61
    |         |    `--A. nostorica Monostori 1994L05
    |         `--Acratia Delo 1939ZL87
    |              |--*A. typica Delo 1930BB61
    |              |--A. bicornis Shi 1987ZL87
    |              |--A. comisa Zhang in Zhang & Liang 1987ZL87
    |              |--A. deloi (Geis 1932)ZL87
    |              |--A. oliviferaCB86
    |              |--A. pulchra Posner 1951CB86
    |              `--A. siratchoica Egorov 1953CB86
    `--Cypridoidea [Cypracea, Cypridacea]M03
         |  i. s.: Camdenidea Swain 1953BB61
         |           `--*C. camdenensis Swain 1953BB61
         |         Haworthina Kellett 1935BB61
         |           `--*H. bulleta (Harris & Lalicker 1932) [=Bairdia bulleta]BB61
         |         Hilboldtina Scott & Summerson 1943BB61
         |           `--*H. multiplicata Scott & Summerson 1943BB61
         |         Palaeocypris Brongniart 1876BB61
         |           `--*P. edwardsii Brongniart 1876BB61
         |         Pruvostina Scott & Summerson 1943BB61
         |           `--*P. wanlessi Scott & Summerson 1943BB61
         |         Ranapeltis Bassler 1941BB61
         |           `--*R. typicalis Bassler 1941BB61
         |         Scabriculocypris Anderson 1940BB61
         |           `--*S. trapezoides Anderson 1940BB61
         |         SuchonellinaBB61
         |--Notodromadidae [Notodromadinae, Notodromides, Notodrominae, Notodromini]MD01
         |    |--Notodromas Liljeborg 1853BB61
         |    |    `--*N. monachus (Müller 1776)BB61 [=Cypris monachaBB61, Monoculus monachusL02]
         |    |--Newnhamia King 1855BB61
         |    |    `--*N. fenestrata King 1855BB61
         |    `--Cyprois Zenker 1854BB61
         |         |--*C. marginata (Strauss 1821) [=Cypris marginata]BB61
         |         |--C. ephraimensis Swain 1964WSB93
         |         `--C. tantouensisNG13

*Macrocypria sarsi (Müller 1912) [=Macrocypris sarsi, Bairdia angusta Sars 1866 non Cythere (Bairdia) angusta Jones 1850]BB61

Pontocypris mytiloides (Norman 1862) [=Cythere (Bairdia) mytiloides; incl. Cypris serrulata Sars 1863, *Pontocypris serrulata]BB61

*Type species of generic name indicated


[BB61] Benson, R. H., J. M. Berdan, W. A. van den Bold, T. Hanai, I. Hessland, H. V. Howe, R. V. Kesling, S. A. Levinson, R. A. Reyment, R. C. Moore, H. W. Scott, R. H. Shaver, I. G. Sohn, L. E. Stover, F. M. Swain & P. C. Sylvester-Bradley. 1961. Systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q99–Q421. Geological Society of America and University of Kansas Press.

[CC02] Cabral, M. C., & J.-P. Colin. 2002. Taxonomie et paléoécologie de nouveaux ostracodes limniques Candonidae dans l’Oxfordien (Jurassique supérieur) du Portugal. Geodiversitas 24: 61–76.

[C01] Chapman, F. 1901. On some fossils of Wenlock age from Mulde, near Klinteberg, Gotland; with notes by Prof. T. Rupert Jones and Dr. F. A. Bather. Annals and Magazine of Natural History, series 7, 7: 141–160, pl. 3.

[CB86] Chen D.-Q. & Bao H. 1986. Lower Permian ostracodes from the Chihsia Formation of Jurong and Longtan, Jiangsu Province. Acta Micropalaeontologica Sinica 3 (2): 107–132.

[F08] Fuhrmann, R. 2008. Die Ostrakoden- und Molluskenfauna des Auelehmprofils Zeitz (Landkreis Burgenland) und ihre Aussage zum Klima sowie zur Landnutzung im jüngeren Holozän Mitteldeutschlands. Mauritiana 20 (2): 253–281.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 4. Familles naturelles des genres. F. Dufart: Paris.

[L05] Liebau, A. 2005. A revised classification of the higher taxa of the Ostracoda (Crustacea). Hydrobiologia 538: 115–137.

[M03] Martens, K. 2003. On a remarkable South African giant ostracod (Crustacea, Ostracoda, Cyprididae) from temporary pools, with additional appendages. Hydrobiologia 500: 115–130.

[MD01] Martin, J. W., & G. E. Davis. 2001. An updated classification of the Recent Crustacea. Natural History Museum Los Angeles County, Science Series 39: 1–124.

[M61] Moore, R. C. 1961. Supplement—USSR treatise on Ostracoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q422–Q429. Geological Society of America and University of Kansas Press.

[NG13] Ni, X., D. L. Gebo, M. Dagosto, J. Meng, P. Tafforeau, J. J. Flynn & K. C. Beard. 2013. The oldest known primate skeleton and early haplorhine evolution. Nature 498: 60–64.

[WSB93] Whatley, R. C., D. J. Siveter & I. D. Boomer. 1993. Arthropoda (Crustacea: Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 343–356. Chapman & Hall: London.

[ZL87] Zhang X.-J. & Liang X.-Y. 1987. Ostracoda from the Taiyuan Formation of Xingyang and Gong Xian districts, Henan. Acta Micropalaeontologica Sinica 4 (3): 293–312.

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