Roquettea singularis, copyright A. Anker.

Belongs within: Cosmetidae.

The Discosomaticinae are cosmetid harvestmen with pectinate claws on the hind two pairs of legs (Medrano et al. 2022).

Characters (from Medrano et al. 2022): Lateral borders of dorsal scutum following general body curvature. Pattern of white blots variable, mostly reticulated or punctate in Roquetteini, with a blurred ring in some Sibambea. Coxa IV usually covered by dorsal sutum so that only posterior margin is apparent. Tarsomeres of leg III seven or more, often nine. Tarsal claws of legs III–IV bipectinated (provided with two rows of denticles). Lateral patches of cteniform T4 microsetae restricted to corners of ventral plate. Wattle stylar barbs reaching ventral margin of stylus and forming a ventral peak.

<==Discosomaticinae [Discosominae, Protinae]MKM22
    |    |--Marronia Medrano, Kury & Mendes 2022MKM22
    |    |    `--*M. elegans (Roewer 1947) [=Metagryne elegans]MKM22
    |    `--+--Paraprotus Roewer 1912MKM22, K03
    |       |    |--*P. atroluteus Roewer 1912K03
    |       |    `--P. albithorax Roewer 1947K03
    |       `--+--Sibambea Roewer 1917MKM22, K03 (see below for synonymy)
    |          |    |--S. cincta (Perty 1833)MKM22, K03 (see below for synonymy)
    |          |    `--+--*S. rotunda Roewer 1917K03, MKM22, K03
    |          |       `--S. waorani Medrano, Kury & Mendes 2022MKM22
    |          `--+--Kayania Medrano, Kury & Mendes 2022MKM22
    |             |    |--*K. bicolumnata (Roewer 1959) [=Metagryne bicolumnata]MKM22
    |             |    `--K. compressicoxa (Avram & Soares 1983) [=Cynorta compressicoxa]MKM22
    |             `--Protus Simon 1879MKM22, K03
    |                  |--P. distinctus (Avram & Soares 1983) [=Discosomaticus distinctus]MKM22
    |                  `--+--P. bolivari (Roewer 1952) [=Cynortula bolivari]MKM22
    |                     `--+--P. marllusi Medrano, Kury & Mendes 2022MKM22
    |                        `--+--*P. insolens Simon 1879K03, MKM22, K03
    |                           |--P. quadripunctatus (Roewer 1947) [=Paraprotus quadripunctatus]MKM22
    |                           `--P. speciosus (Roewer 1928) [=Paraprotus speciosus]MKM22
         |--Gryne Simon 1879MKM22, K03
         |    |  i. s.: G. marmorata (Roewer 1915) [=Poecilaema marmoratum, Paecilaema marmoratum]MKM22
         |    |--G. marginalis (Perty 1833)MKM22, K03 (see below for synonymy)
         |    `--+--G. dimorpha Mello-Leitão 1928MKM22, K03
         |       `--+--G. coccinelloides (Mello-Leitão 1935)MKM22, K03 (see below for synonymy)
         |          |--G. leprosa Sørensen in Henriksen 1932MKM22 (see below for synonymy)
         |          `--G. orensis (Sørensen 1879)MKM22, K03 (see below for synonymy)
         `--Roquettea Mello-Leitão 1931 [incl. Tetracyphus Sørensen in Henriksen 1932 non Chevrolat 1881]MKM22
              |--R. andina (Roewer 1947) [=Gryne andina]MKM22
              |--R. jalapensis Ferreira & Kury 2010MKM22
              `--+--R. taurina Ferreira & Kury 2010MKM22
                 `--+--R. peba Kury & Ferreira 2012MKM22
                    `--+--+--R. bubalina Medrano & Kury 2018MKM22
                       |  `--R. troguloides Medrano & Kury 2018MKM22
                       `--+--R. carajas Kury & Ferreira 2012MKM22
                          `--+--R. decioi Kury 2013MKM22
                             `--+--*R. singularis Mello-Leitão 1931K03, MKM22, K03 [incl. *Tetracyphus mirabilis Sørensen 1932K03]
                                `--R. scrotalis Ferreira & Kury 2010MKM22

Gryne coccinelloides (Mello-Leitão 1935)MKM22, K03 [=Poecilaema coccinelloidesK03; incl. Gryne reticulata Piza 1938K03, Iryne (l. c.) reticulataK03]

Gryne leprosa Sørensen in Henriksen 1932MKM22 [=Poecilaema leprosum msK03; incl. G. perlata Mello-Leitão 1936MKM22, G. vermiculata Mello-Leitão 1944MKM22]

Gryne marginalis (Perty 1833)MKM22, K03 [=Cosmetus marginalisK03, Paecilaema marginale non Cynorta marginalis Banks 1909K03, Poecilaema marginaleK03; incl. Gryne amazonica Roewer 1947MKM22, *G. paraensis Simon 1879K03]

Gryne orensis (Sørensen 1879)MKM22, K03 [=Cosmetus orensisK03; incl. G. angustipes Mello-Leitão 1931K03, G. angusticeps (l. c.)K03, Paecilaema bonita msK03, P. neglectum Ringuelet 1957MKM22]

Sibambea Roewer 1917MKM22, K03 [incl. Discosoma Perty 1833 non Rueppell & Leuckart 1828K03, Discosomaticus Roewer 1923MKM22, Discosominana Strand 1942K03]

Sibambea cincta (Perty 1833)MKM22, K03 [=*Discosoma cinctumK03, Cosmetus cinctusK03, *Discosomaticus cinctusK03, *Discosominana cinctaK03; incl. Discosomaticus sturmi Roewer 1963MKM22]

*Type species of generic name indicated


[K03] Kury, A. B. 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Ibérica de Aracnología, volumen especial monográfico 1: 1–337.

[MKM22] Medrano, M., A. B. Kury & A. C. Mendes. 2022. Morphology-based cladistics splinters the century-old dichotomy of the pied harvestmen (Arachnida: Gonyleptoidea: Cosmetidae). Zoological Journal of the Linnean Society 195 (2): 585–672.

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