Bandidus breviusculus, copyright Dhobern.

Belongs within: Myrmeleontidae.

The Distoleontini are a group of antlions with relatively short cubital veins in the forewing (New 1985a).

Characters (from New 1985a): Forewing costal cells simple; veins CuA2 and (CuP + 1A) relatively short; vein 2A close to 1A at base, then 2A and 3A closely associated over central lengths. Hindwing Rs arising well before medial fork; one (rarely two or more) presectoral crossveins.

<==Distoleontini [Formicaleonini, Formicaleontini, Glenurini, Macronemurinae, Macronemurini]
    |--NeuroleonVM20 [incl. Neleomma Kimmins 1938 nec Navás 1914N85b]
    `--+--Macronemurus Costa 1835VM20, N85b
       |    |--*M. appendiculatusN85b
       |    `--M. striola Kolbe 1897HM55
       `--+--Euroleon nostrasVM20
          `--Distoleon Banks 1910VM20, N85b [incl. Eidoleon Esben-Petersen 1918N85b]
               |--D. somnolentus (Gerstaecker 1884) (see below for synonymy)N85b
               |--D. bistrigatus (Rambur 1842) (see below for synonymy)N85b
               |--D. morpheus (Kirby 1900) [=Formicaleo morpheus]N85b
               |--D. nefarius Navás 1910 (n. d.)N85b
               |--D. perjurusN85b
               `--D. tetragrammicusVM20
Distoleontini incertae sedis:
  ‘Formicaleon’ audaxN85b
  Brachyleon Tillyard 1916N85b
    `--*B. darwini (Banks 1915) [=Macronemurus darwini]N85b
  Eophanes Banks 1931N85b
    |--*E. formosaN85b
    `--E. distinctus (Banks 1939) [=Formicaleo distinctus; incl. E. falcata Kimmins 1955]N85b
  Stenogymnocnemia Esben-Petersen 1923N85b
    |--*S. malus (Walker 1853) [=Myrmeleon malus, Glenurus malus]N85b
    `--S. angusta New 1985N85b
  Xantholeon Tillyard 1916 [incl. Cataleon Navás 1933]N85b
    |--*X. helmsi Tillyard 1916 [incl. *Cataleon gratiosus Navás 1933]N85b
    |--X. lineatus New 1985N85b
    |--X. manselli New 1985N85b
    |--X. montanus New 1985N85b
    `--X. xadnus New 1985N85b
  Stenoleon Tillyard 1916N85b
    |--*S. fieldi Tillyard 1916N85b
    |--S. cingulatus New 1985N85b
    |--S. copleyensis New 1985N85b
    |--S. gradostriatus New 1985N85b
    |--S. grandithecus New 1985N85b
    `--S. navasi New 1985N85b
  Escura Navás 1914 [incl. Alloformicaleon Esben-Petersen 1915]N85b
    |--*E. divergens Navás 1914N85b
    |--E. australis (Esben-Petersen 1915) [=Formicaleon australis, *Alloformicaleon australis]N85b
    |--E. fitzroyensis New 1985N85b
    |--E. josephinae New 1985N85b
    |--E. nigrosignatus (Tillyard 1916) [=Distoleon nigrosignatus, Eidoleon nigrosignatus]N85b
    |--E. notostriatus New 1985N85b
    |--E. patriciae New 1985N85b
    |--E. punctata New 1985N85b
    `--E. rhondae New 1985N85b
  Creoleon nubifer (Kolbe 1897)N85a, HM55 [=Creagris nubiferHM55, Gama nubiferHM55]
  Capophanes Banks 1938B38
    `--*C. conspersa Banks 1938B38
  Bandidus Navás 1914 [incl. Heteroleon Esben-Petersen 1918]N85b
    |--*B. canifrons (Navás 1914) [=Formicaleo canifrons, Alloformicaleon canifrons]N85b
    |--B. abditus (Walker 1853) [=Myrmeleon abditus, Macronemurus abditus]N85b
    |--B. aeratus New 1985N85b
    |--B. amarillus New 1985N85b
    |--B. antestriatus New 1985N85b
    |--B. apicalis (Esben-Petersen 1923) [=Heteroleon apicalis]N85b
    |--B. barradalensis New 1985N85b
    |--B. breviusculus (Gerstaecker 1885) (see below for synonymy)N85b
    |--B. brisbanensis New 1985N85b
    |--B. centralis New 1985N85b
    |--B. congestus (Gerstaecker 1885) [=Myrmeleon congestus, Formicaleo congestus]N85b
    |--B. cornutus New 1985N85b
    |--B. dalyensis New 1985N85b
    |--B. dispersus (Banks 1910) [=Formicaleo dispersus]N85b
    |--B. exilis (Esben-Petersen 1918) [=Heteroleon exilis]N85b
    |--B. furcatus New 1985N85b
    |--B. fuscus New 1985N85b
    |--B. handschini New 1985N85b
    |--B. hilli (Esben-Petersen 1923) [=Formicaleo hilli]N85b
    |--B. hyalinus (Tillyard 1916) [=Alloformicaleon hyalinus, Formicaleo hyalinus]N85b
    |--B. kimminsi New 1985N85b
    |--B. longigona New 1985N85b
    |--B. marginalis (Banks 1910) [=Formicaleo marginalis, Heteroleon marginalis]N85b
    |--B. nebulosus New 1985N85b
    |--B. nigrifrons (Navás 1922) [=Nelees nigrifrons, Neleomma nigrifrons]N85b
    |--B. occidentalis New 1985N85b
    |--B. ocellonotus New 1985N85b
    |--B. oppositus New 1985N85b
    |--B. pulchellus (Esben-Petersen 1923) [=Heteroleon pulchellus]N85b
    |--B. rydalmerensis New 1985N85b
    |--B. septus (Gerstaecker 1885) [=Formicaleon septus]N85b
    |--B. spinosus New 1985N85b
    |--B. tibooburranensis New 1985N85b
    |--B. umbronotus New 1985N85b
    |--B. vafer (Walker 1853) (see below for synonymy)N85b
    `--B. xylotus New 1985N85b

Bandidus breviusculus (Gerstaecker 1885) [=Myrmeleon breviusculus, Formicaleo breviusculus; incl. Alloformicaleon waterhousei Tillyard 1916]N85b

Bandidus vafer (Walker 1853) [=Myrmeleon vafer, Formicaleo vafer; incl. Myrmeleon desperatus Walker 1853, Formicaleon desperatus, Macronemurus desperatus, Myrmeleon malefidus Walker 1853, Myrmecaelurus malefidus, Myrmeleon perniciosus Walker 1853, Formicaleo perniciosus, Macronemurus perniciosus, Nelees strigatus Navás 1914]N85b

Distoleon bistrigatus (Rambur 1842) [=Myrmeleon bistrigatus, Eidoleon bistrigatus; incl. Formicaleo acuminatus Okamoto 1910, F. brahmanicus Banks 1913, Myrmeleon striola Walker 1853, F. striola, M. torvus Walker 1853, F. torvus]N85b

Distoleon somnolentus (Gerstaecker 1884) [=Formicaleo somnolentus; incl. D. peterseni Navás 1935, F. thoreyi Navás 1914, *D. verticalis Banks 1910]N85b

*Type species of generic name indicated


[B38] Banks, N. 1938. Two new genera of Myrmeleonidae. Proceedings of the Entomological Society of Washington 40 (5): 125–129.

[HM55] Handschin, E. & W. Markl. 1955. Neuropteren aus Angola. Diamang Publicações Culturais 27: 65–82.

[N85a] New, T. R. 1985a. A revision of the Australian Myrmeleontidae (Insecta: Neuroptera). I. Introduction, Myrmeleontini, Protoplectrini. Australian Journal of Zoology, Supplementary Series 104: 1–90.

[N85b] New, T. R. 1985b. A revision of the Australian Myrmeleontidae (Insecta: Neuroptera). III. Distoleontini and Acanthaclisinae. Australian Journal of Zoology, Supplementary Series 106: 1–159.

[VM20] Vasilikopoulos, A., B. Misof, K. Meusemann, D. Lieberz, T. Flouri, R. G. Beutel, O. Niehuis, T. Wappler, J. Rust, R. S. Peters, A. Donath, L. Podsiadlowski, C. Mayer, D. Bartel, A. Böhm, S. Liu, P. Kapli, C. Greve, J. E. Jepson, X. Liu, X. Zhou, H. Aspöck & U. Aspöck. 2020. An integrative phylogenomic approach to elucidate the evolutionary history and divergence times of Neuropterida (Insecta: Holometabola). BMC Evolutionary Biology 20: 64.

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