Siragra sp., copyright Stephane Hanot.

Belongs within: Ichneumonoidea.
Contains: Rogadinae, Braconinae, Opiinae, Alysiinae.

The Doryctinae are a cosmopolitan group of braconid wasps with ectoparasitic larvae, most commonly on concealed Coleoptera larvae. The Neotropical Ypsistocerini, distinguished from other tribes by an enlarged telotarsus, are known from termite nests but their biology is otherwise unknown (Achterberg 1993).

Characters (from Achterberg 1993): Maxillary palp with six segments; hypoclypeal depression deep and wide, middle of apparent ventral margin of clypeus distinctly above upper level of mandibular bases, labrum concave; eyes wider than base of mandible; posterior flange of propleuron present, largely dorsally situated; fore tibia often with row of stout pegs or spines, at most as long as about six times their length; hind coxa often angulate anteroventrally, frequently with a ventral tubercle; ovipositor dorsally nearly always with double nodus subapically.

    |--+--+--Liobracon aquiloniusBS23
    |  |  `--Odontobracon montanusBS23
    |  `--+--EcphylusBS23
    |     |    |--E. oculatus Muesebeck 1960P92
    |     |    `--E. rohweriBS23
    |     `--+--Bolivar ornaticornisBS23
    |        `--+--Notiospathius terminalisBS23
    |           `--+--+--Pioscelus borealisBS23
    |              |  `--HeterospilusBS23
    |              |       |--H. microstigmiBM76
    |              |       `--H. prosopidisN57
    |              `--+--Coiba jeffersoniBS23
    |                 `--+--Leluthia mexicanaBS23
    |                    `--Callihormius bifasciatusBS23
       |  |  |    |--Spathioplites phreneticusA93
       |  |  |    `--SpathiusBS23
       |  |  |         |--S. clavatus [=Bracon (Spathius) clavatus]G20
       |  |  |         `--S. laflammeiBS23
       |  |  `--Rhaconotus fasciatusBS23
       |  `--+--PambolinaeA93
       |     |    |--Notiopambolus depressicaudaA93
       |     |    `--Pambolus biglumisBS23, A93
       |     `--+--OntsiraBS23
       |        `--DoryctiniA93
       |             |--Priosphys biannulatusA93
       |             |--Doryctes erythromelasHR11
       |             `--Dendrosoter Wesmael 1838A93, M38 [incl. Caenopachys Foerster 1862 M38, Eurybolus Ratzeburg 1848M38]
       |                  |--*D. protuberans [=Bracon protuberans; incl. B. curtisii, *Eurybolus curtisii]M38
       |                  |--D. hartigii [=Bracon hartigii, *Caenopachys hartigii]M38
       |                  |--D. integer Muesebeck 1938M38
       |                  |--D. middendorffiiM38
       |                  |--D. scaber Muesebeck 1938M38
       |                  |--D. scolytivorus (Viereck & Rohwer 1913) [=Caenopachys scolytivora]M38
       |                  `--D. sulcatus Muesebeck 1938M38
       `--+--+--Rhysipolinae [Rhysipolini]BS23
          |  |    |--RhysipolisA93
          |  |    |    |--R. hariolatorA93
          |  |    |    `--R. meditatorA93
          |  |    `--CantharoctonusBS23
          |  `--+--RogadinaeBS23
          |     `--+--BetylobraconinaeA93
          |        |    |--Mesocentrus crassipesA93
          |        |    `--Betylobracon waterhouseiA93
          |        `--Mesostoa [Mesostoinae]A93
          |             |--M. compressaA93
          |             `--M. kerriA93
             |    |--GnamptodonBS23
             |    |    |--G. molestusA93
             |    |    `--G. pumilioA93
             |    `--PseudognaptodonBS23
             `--+--+--Vaepellis [Vaepellinae]A93
                |  |    `--V. varicaA93
                |  `--+--BraconinaeBS23
                |     `--Telengaia [Telengaiinae]A93
                |          `--T. ventralisA93
                `--+--Exothecinae [Exothecini]A93
                   |    |--Xenarcha abnormisA93
                   |    `--ColastesBS23
                   |         |--C. braconiusA93
                   |         `--C. hariolatorMS01
Doryctinae incertae sedis:
  Stenocorse Marsh 1968A93
  ‘Atanycolus’ tomentosusQI93
  Termitobracon [Ypsistocerinae, Ypsistocerini]A93
    `--T. emersoniA93
  Evaniodes [Evaniodini]A93
    `--E. areolatusA93
  Pareucorystes [Hecabolini]A93
    `--P. varinervisA93
  Leptorhaconotus [Leptorhaconotini]A93
    `--L. brunneusA93
    `--Siragra nitidaA93

*Type species of generic name indicated


[A93] Achterberg, C. van. 1993. Illustrated key to the subfamilies of the Braconidae (Hymenoptera: Ichneumonoidea). Zoologische Verhandelingen 283: 1–189.

[BS23] Blaimer, B. B., B. F. Santos, A. Cruaud, M. W. Gates, R. R. Kula, I. Mikó, J.-Y. Rasplus, D. R. Smith, E. J. Talamas, S. G. Brady & M. L. Buffington. 2023. Key innovations and the diversification of Hymenoptera. Nature Communications 14: 1212.

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[HR11] Heraty, J., F. Ronquist, J. M. Carpenter, D. Hawks, S. Schulmeister, A. P. Dowling, D. Murray, J. Munro, W. C. Wheeler, N. Schiff & M. Sharkey. 2011. Evolution of the hymenopteran megaradiation. Molecular Phylogenetics and Evolution 60: 73–88.

[M38] Muesebeck, C. F. W. 1938. The genus Dendrosoter Wesmael in the United States (Hymenoptera: Braconidae). Proceedings of the Entomological Society of Washington 40 (9): 281–287.

[N91] Naumann, I. D. 1991. Hymenoptera (wasps, bees, ants, sawflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 2 pp. 916–1000. Melbourne University Press: Carlton (Victoria).

[N57] Nicholson, A. J. 1957. The self-adjustment of populations to change. Cold Spring Harbor Symposia on Quantitative Biology 22: 153–172.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[QI93] Quicke, D. L. J., & S. N. Ingram. 1993. Braconine wasps of Australia. Memoirs of the Queensland Museum 33 (1): 299–336.

[R00] Ross, E. S. 2000. Embia: Contributions to the biosystematics of the insect order Embiidina. Occasional Papers of the California Academy of Sciences 149: 1–53, 1–36.

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