Ectopsocus briggsi, copyright Christophe Quintin.

Belongs within: Lachesilloidea.

Ectopsocus is a genus of barklice including some of the most widespread psocid species not directly associated with humans. Males have a clunium bearing a row of teeth or other ornamentation; females have all three pairs of gonapophyses present and the subgenital plate is usually bilobed and setose (New & Lienhard 2007). Wings have dark spots at the ends of veins in the cosmopolitan species E. briggsi and E. californicus (Schmidt & New 2008).

<==Ectopsocus McLachlan 1899SN08 (see below for synonymy)
    |--*E. briggsi McLachlan 1899SN08 [incl. E. limbatus Navás 1909L98]
    |--E. adelphos Thornton & Wong 1968NL07
    |--E. aethiops (Hagen 1859)NL07
    |--E. amblyura Thornton & Wong 1968NL07
    |--E. amphithrix Thornton & Wong 1968NL07
    |--E. anisodontus Li 2002L02
    |--E. argus Thornton & Wong 1968NL07
    |--E. axillaris (Smithers 1969) [=Interpsocus axillaris]SN08
    |--E. baliosus Thornton & Wong 1968NL07
    |--E. barlowi New & Lee 1992NL07
    |--E. basalis (Banks 1937)NL07
    |--E. berlesii Ribaga 1900 (n. d.)L98
    |--E. bicaudatusB55
    |--E. bifurcatus Vaughan, Thornton & New 1991NL07
    |--E. brinchangensis New & Lee 1992NL07
    |--E. brunneus (Edwards 1950)SN08 [=*Interpsocus brunneusNL07]
    |--E. californicus (Banks 1903)SN08 (see below for synonymy)
    |--E. cetratusSN08
    |--E. cinctus Thornton 1962 [=Estipulaceus cinctus]NL07
    |--E. cirratus Thornton & Wong 1968NL07
    |--E. cluniatus Vaughan, Thornton & New 1989NL07
    |--E. coccophilus Ball 1943NL07
    |--E. columbianus Badonnel 1986B86
    |--E. comitus Thornton & Wong 1968 [=Estipulaceus comitus]NL07
    |--E. coyae Schmidt & New 2008SN08
    |--E. crinitus Thornton & Wong 1968NL07
    |--E. cristatus Thornton & Wong 1968NL07
    |--E. decenipunctatus Li 1993 [=E. decemipunctatus (l. c.), E. decenipnctatus]L03
    |--E. denervus Thornton & Wong 1968NL07
    |--E. denotatus Thornton & Wong 1968NL07
    |--E. dicroglossus Thornton & Wong 1968NL07
    |--E. drepanus Thornton 1984NL07
    |--E. edwardsi New 1973SN08
    |--E. equidentus Li 1999 [=E. equicornutus]L03
    |--E. erosus (Enderlein 1903)NL07
    |--E. ewarti Vaughan, Thornton & New 1989NL07
    |--E. femestratus Thornton & Wong 1968L02
    |--E. fumidus Thornton & Wong 1968NL07
    |--E. furcatus Thornton & Wong 1968NL07
    |--E. gentingensis New & Lee 1992NL07
    |--E. ghesquierei Ball 1943L02
    |--E. graminus Schmidt & New 2008SN08
    |--E. gyroancistrus Li 2002L02
    |--E. heurni (Navás 1924)NL07
    |--E. hickmani Schmidt & New 2008SN08
    |--E. hirsutus Thornton 1962NL07 [=Ectotrichus hirsutusL03]
    |--E. hypandrus Thornton 1984NL07
    |--E. inclinans Li 2002L02
    |--E. innotatus Thornton & Wong 1968NL07
    |--E. insularis Smithers & Thornton 1974L02
    |--E. intersitus Thornton & Wong 1968NL07
    |--E. isodentus Li 1997L02
    |--E. kepongensis New & Lee 1992NL07
    |--E. leteusistrius Li 2002L02
    |--E. longitudinalis Li 2002L02
    |--E. machadoi Badonnel 1955B55
    |--E. macrotaenious Li 2002L02
    |--E. maindroni Badonnel 1935 [=Estipulaceus maindroni]NL07
    |--E. marginatusM93
    |--E. meridionalis Ribaga 1904NL07 [=E. briggsi var. meridionalisL98]
    |--E. musae (Kunstler & Chaine 1902) (n. d.)L98
    |--E. myrmecophilus (Enderlein 1903)NL07
    |--E. nerens (Hickman 1934) [=Micropsocus nerens]SN08
    |--E. nidicolus Thornton & Wong 1968NL07
    |--E. nyingchiensis Li 2002 [=E. nyingchinensis]L02
    |--E. ornatus Thornton 1962NL07 [=Ectotrichus ornatusL03]
    |--E. paraplesius Thornton & Wong 1968NL07
    |--E. pearmaniMG56
    |--E. petersi Smithers 1978 [incl. E. australis Schmidt & Thornton 1993]SN08
    |--E. pictus Mockford 1974M74
    |--E. pilosus Badonnel 1967NL07
    |--E. predanus Vaughan, Thornton & New 1989NL07
    |--E. proctus New & Lee 1992NL07
    |--E. punctatusM83
    |--E. quadratiapicius Li 2002 [=E. quadratiapicis]L03
    |--E. quadrisetus Li 2002L02
    |--E. richardsi (Pearman 1929)L03 [=*Chaetopsocus richardsiNL07, Ectotrichus richardsiL03]
    |--E. ridderi Vaughan, Thornton & New 1989NL07
    |--E. rileyae Schmidt & Thornton 1993SN08
    |--E. risdonensis Schmidt & New 2008SN08
    |--E. salpinx Thornton & Wong 1968NL07
    |--E. setulosus Li 1997L03, L02 [=*Ectotrichus setulosusL03]
    |--E. similis Badonnel 1955B55
    |--E. speciosus Thornton & Wong 1968NL07
    |--E. spilocephalus Li 1993L02
    |--E. spilotusM93
    |--E. sprenti Schmidt & New 2008SN08
    |--E. stictus Thornton & Wong 1968NL07
    |--E. strauchi Enderlein 1906 [incl. Peripsocus opulentus Navás 1908]L98
    |--E. strictifoliatus Li 2002L02
    |--E. sumatrensis Vaughan, Thornton & New 1991NL07
    |--E. tenellus Thornton & Wong 1968NL07
    |--E. thibaudi Badonnel 1979M93
    |--E. thoi New & Lee 1992NL07
    |--E. thysanus Thornton & Wong 1968NL07
    |--E. tinctus Navás 1924NL07
    |--E. titschacki Jentsch 1939NL07 [incl. E. gabelensis Badonnel 1955L98]
    |--E. triangulus Thornton & Wong 1968NL07
    |--E. tridentatus Thornton 1962L02 [=E. meridionalis tridentatusL98]
    |--E. uncinatusM93
    |--E. vachoni Badonnel 1945NL07 [incl. E. dimorphus Mockford & Gurney 1956L98]
    |--E. vacuus New & Lee 1992NL07
    |--E. vannus Thornton & Wong 1968NL07
    |--E. venosus Lienhard 2002NL07
    |--E. vilhenai Badonnel 1955NL07
    |--E. vishnyakovae Schmidt 1993 (see below for synonymy)L98
    |--E. waterstradti (Enderlein 1901) [=*Micropsocus waterstradti]NL07
    |--E. yongi New & Lee 1992NL07
    `--E. zayuensis Li 2002L02

Ectopsocus McLachlan 1899SN08 [incl. Chaetopsocus Pearman 1929L03, Ectotrichus Li 2002L03, Interpsocus Edward 1950L02, Micropsocus Enderlein 1901L02]

Ectopsocus californicus (Banks 1903)SN08 [=Peripsocus californicusSN08; incl. E. congener Tillyard 1923M93, E. froggatti Enderlein 1906SN08]

Ectopsocus vishnyakovae Schmidt 1993 [incl. E. brunneus Vishnyakova in Svadzhyan et al. 1963 non Interpsocus brunneus Edwards 1950]L98

*Type species of generic name indicated


[B55] Badonnel, A. 1955. Psocoptères de l’Angola. Diamang Publicações Culturais 26: 11–267.

[B86] Badonnel, A. 1986. Psocoptères de Colombie (Insecta, Psocoptera). Spixiana 9 (2): 179–223.

[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[L98] Lienhard, C. 1998. Faune de France. France et Régions Limitrophes. 83. Psocoptères Euro-Méditerranéens. Fédération Française des Sociétés de Sciences Naturelles: Paris.

[L03] Lienhard, C. 2003. Nomenclatural amendments concerning Chinese Psocoptera (Insecta), with remarks on species richness. Revue Suisse de Zoologie 110 (4): 695–721.

[M83] Martin, N. A. 1983. Miscellaneous observations on a pasture fauna: an annotated species list. DSIR Entomology Division Report 3: 1–98.

[M74] Mockford, E. L. 1974. Records and descriptions of Cuban Psocoptera. Entomologica Americana 48 (2): 103–215.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[MG56] Mockford, E. L., & A. B. Gurney. 1956. A review of the psocids, or book-lice and bark-lice, of Texas (Psocoptera). Journal of the Washington Academy of Sciences 46 (11): 353–368.

[NL07] New, T. R., & C. Lienhard. 2007. The Psocoptera of Tropical South-east Asia. Brill: Leiden.

[SN08] Schmidt, E. R., & T. R. New. 2008. The Psocoptera (Insecta) of Tasmania, Australia. Memoirs of Museum Victoria 65: 71–152.

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