Priopus castaneus, copyright Terrier3538.

Belongs within: Elateridae.
Contains: Glyphochilus, Melanoxanthus, Synaptini, Agriotini, Ampedini, Megapenthini, Dicrepidiini, Elaterini, Melanotini, Cebrionini, Physorhinini.

The Elaterinae are a group of click beetles in which the head is distinctly convex anteriorly, and the wings bear a set of three apical sclerotisations (Calder 1996). The Aplastini have a long prosternum and short, indistinct tibial spurs (Johnson 2002).

The Adrastini
Published 21 March 2018
Glyphonyx sp., copyright Mike Quinn.

For the subject of my next post, I drew the click beetle tribe Adrastini. Well, actually, I drew the tribe Synaptini but dibs on that name was originally called by a family of sea cucumbers so it seems that ‘Adrastini’ should be the tribe’s proper name.

The Adrastini are one of those groups of animals for which there seems to be little known to discuss other than their general morphology. The more typical click beetles tend to be fairly uniform in their general appearance and are often not that easy to distinguish. Adrastins belong to the subfamily Elaterinae and resemble other elaterines in their deflexed mouthparts, arcuate prosternum, open mesocoxal cavities and tarsal claws without basal setae. They differ from other elaterines in having serrate tarsal claws according to Stibick (1979). Mind you, another related group, the Melanotini, are supposed to be distinguished by their pectinate claws and I’m not entirely sure what the difference between ‘serrate’ and ‘pectinate’ is supposed to be in this context.

The Adrastini are widespread though they seem to be absent from Australia. Several genera are recognised; one of these, Glyphonyx, is distinctly larger than the rest and includes about half the tribe’s known species. As far as I know, the larva has never been described for any member of this group, so what they are doing ecologically remains a largely unknown quantity.

Systematics of Elaterinae

Characters (from Calder 1996): Head distinctly convex anteriorly. Frontal carina usually complete across front between eyes or incomplete medially. Antennae nearly always serrate or subserrate, rarely pectinate. Prosternal spine (posterior to procoxae) usually longer than procoxal diameter, rarely shorter than procoxal diameter or same length as procoxal diameter. Mesocoxal cavity open to both mesepimeron and mesepisternum, rarely open to mesepimeron only. Meso-trochantin visible, rarely not visible. Hind wing with short radial cell; vein MP4 with apparent cross vein to CuA2, rarely without; wedge cell present, rarely absent; apex of wing membrane (without venation) occupying less than 0.2× length of wing membrane; apex with three sclerotisations arranged in form of Greek letter ψ. Tarsal claws without basal setae on outer flat portion.

<==Elaterinae (see below for synonymy)KB11
    |--+--Ludioschema yushiroi Suzuki 1999KB11, PS07 [=Agonischius obscuripes var. sanguinicollis Miwa 1928PS07]
    |  `--+--AgriotiniKB11
    |     `--AdrastusE12 [AdrastiniKB11]
    |          `--A. bicolor Lucas 1846E12
    |--Aplastini [Aplastinae, Plastoceridae]KB11
    |    |--Aplastus LeConte 1859 [incl. Anamesus LeConte 1866]J02
    |    |--Euthysanius LeConte 1853J02
    |    `--Plastocerus LeConte 1853 [incl. Octinodes Candèze 1863]J02
    |         |--P. angulosus (Germar 1845)B14
    |         `--P. thoracicusB14
       |  |  `--+--MegapenthiniKB11
       |  |     `--DicrepidiiniKB11
       |  `--+--TomicephalusKB11
       |     `--PriopusKB11
       |          |--P. angulatus (Candèze 1860)PS07
       |          |--P. castaneus (Miwa 1930) [=Neodiploconus castaneus]PS07
       |          |--P. humeralisKB11
       |          |--P. ornatusKB11
       |          `--P. rufulus (Candèze 1891)PS07
          |  `--MelanotiniKB11
Elaterinae incertae sedis:
  Ophidius Candèze 1863C96
    |--*O. elegans Candèze 1863C96
    |--O. dracunculus Candèze 1863C96
    |--O. histrio (Boisduval 1835)B14 [=Elater histrioC96, Agrypnus histrioB35]
    `--O. vericulatus Neboiss 1975C96
  Anilicus Candèze 1863C96
    |--*A. attenuatus Candèze 1863 [=A. attenatus (l. c.)]C96
    |--A. loricatus Candèze 1863C96
    |--A. nigroterminatus Macleay 1827M86
    |--A. parvus Gullan 1977C96
    |--A. rectilineatus Gullan 1977C96
    `--A. xanthomus (Macleay 1826)C96 (see below for synonymy)
  Acroniopus Erichson 1843 [=Atelopus Erichson 1842 non Duméril & Bibron 1841]C96
    |--*A. humilis (Erichson 1842) [=*Atelopus humilis]C96
    |--A. ater Candèze 1863C96
    |--A. fuliginosus Candèze 1863C96
    |--A. infimus (Erichson 1842) [=Atelopus infimus]C96
    |--A. marginicollis Schwarz 1906C96
    |--A. pallidus Blackburn 1889C96
    |--A. pubescens Macleay 1872C96
    |--A. rufipennis Macleay 1872 [incl. Anilicus flavipennis Candèze 1878]C96
    `--A. sydneyanus (Boisduval 1835) [=Elater sydneyanus]C96
  Anilicoides Candèze 1893C96
    |--*A. depressus Candèze 1893C96
    `--A. haemorrhoidalis (Candèze 1889) [=Anilicus haemorrhoidalis]C96
  Ascesis Candèze 1863C96
    |--*A. australis Candèze 1863C96
    |--A. campyloides Candèze 1897C96
    |--A. impurus (Germar 1848) [=Pristilophus impurus]C96
    |--A. mastersii Macleay 1872C96
    `--A. testaceus Elston 1930C96
  Augenotus Gullan 1977C96
    |--*A. quadriguttatus (Erichson 1842) [=Melanoxanthus quadriguttatus, Anilicus quadriguttatus]C96
    |--A. aurantius Gullan 1977C96
    `--A. australis (Candèze 1859) [=Melanoxanthus australis]C96
  Diadysis Candèze 1882C96
    `--*D. morsi Candèze 1882C96
  Idiomilta Calder 1996C96
    `--*I. queenslandica Calder 1996C96
  Lingana Neboiss 1960C96
    |--*L. illita (Candèze 1863) [=Chrosis illita, Elatichrosis illita; incl. Metablax plasoni Schwarz 1903]N60
    `--L. dombarta Neboiss 1960C96
  Ochosternus Candèze 1863C96
    |--*O. zealandicus [=Elater zealandicus]C96
    |--O. howensis Lea 1929C96
    `--O. norfolcensis Lea 1929C96
  Paranilicus Candèze 1878C96
    |--*P. macleayi Candèze 1878C96
    `--P. brevicornis (Macleay 1872) [=Ophidius brevicornis]C96
  Rangsia Calder 1996C96
    `--*R. macleayi (Candèze 1893) [=Ophidius macleayi]C96
  Simodactylus Candèze 1859C96
    |--*S. cinnamomeus (Boisduval 1835)C96 [=Elater cinnamomeusC96, Aeolus cinnamomeusB35]
    |--S. brunneus Schwarz 1906C96
    `--S. suturalis Schwarz 1906C96
  Yalganus Neboiss 1975C96
    |--*Y. serricornis (Candèze 1878) [=Ophidius serricornis]C96
    `--Y. hirticornis (Carter 1939) [=Dicteniophorus hirticornis]C96

Anilicus xanthomus (Macleay 1826)C96 [=Elater xanthomusC96; incl. An. anticusM86, Ampedus semiflavus Germar 1844C96, An. semiflavusM86]

Elaterinae [Adrastinae, Agriotinae, Ampedidae, Amphilabridae, Athoomorphinae, Cardiorhininae, Cebriognathinae, Cebrionidae, Cebrioninae, Cratonychidae, Dicrepidiinae, Melanotinae, Physorhininae, Steatoderidae]KB11

*Type species of generic name indicated


[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d’Urville. Faune entomologique de l’océan Pacifique, avec l’illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[C96] Calder, A. A. 1996. Click beetles: genera of the Australian Elateridae (Coleoptera). CSIRO Australia: Collingwood.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[J02] Johnson, P. J. 2002. Elateridae Leach 1815. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 160–173. CRC Press.

[KB11] Kundrata, R., & L. Bocak. 2011. The phylogeny and limits of Elateridae (Insecta, Coleoptera): is there a common tendency of click beetles to soft-bodiedness and neoteny? Zoologica Scripta 40: 364–378.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part IV. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 259–380.

[N60] Neboiss, A. 1960. The genera Elatichrosis Hyslop and Lingana, gen. nov. (Coleoptera: Elateridae). Australian Journal of Zoology 8 (2): 289–306.

[PS07] Platia, G., & R. Schimmel. 2007. Click beetles of Taiwan collected by the expeditions of the Hungarian Natural History Museum in the years 1995 to 2003 (Coleoptera: Elateridae). Annales Historico-Naturales Musei Nationalis Hungarici 99: 49–91.

Stibick, J. N. L. 1979. Classification of the Elateridae (Coleoptera). Relationships and classification of the subfamilies and tribes. Pacific Insects 20 (2–3): 145–186.

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