Upland bully Gobiomorphus breviceps, copyright Stella McQ.

Belongs within: Gobiomorpharia.
Contains: Hypseleotris, Eleotris.

The Eleotridae, sleepers, are a clade of often relatively large gobies supported by molecular data.

<==Eleotridae [Eleotrididae, Eleotroidei]
    |--Austrolethops Whitley 1935 [Austrolethopinae]S58
    |    `--*A. wardi Whitley 1935S58
    |    |--Eleotrica Ginsberg 1933S58
    |    |--Chriolepidops Smith 1958S58
    |    |    `--*C. nebulofasciatus Smith 1958S58
    |    |--Satulinus Smith 1958S58
    |    |    `--*S. zanzibarensis Smith 1958S58
    |    |--Dactyleleotris Smith 1958S58
    |    |    `--*D. tentaculatus Smith 1958S58
    |    |--Hetereleotris Bleeker 1874 [incl. Leioeleotris Fowler 1934, Riukiuia Fowler 1946]S58
    |    |    |--*H. diadematus (Ruppell 1828) [=Gobius diadematus, Eleotris diademata, Gobiosoma diadematum]S58
    |    |    |--H. kenyae Smith 1958S58
    |    |    |--‘Riukiuia’ poecila Fowler 1946S58
    |    |    `--H. zonatus (Fowler 1934) [=Leioeleotris zonatus]S58
    |    `--Lioteres Smith 1958S58
    |         |--L. (Lioteres)S58
    |         |    |--*L. (L.) caminatus Smith 1958S58
    |         |    `--L. (L.) vulgare (Klunzinger 1871)S58
    |         `--L. (Pseudolioteres Smith 1958)S58
    |              `--L. (*P.) simulans Smith 1958S58
         |  i. s.: Pogoneleotris microps Weber 1908MWW89, M58 [=Prionobutis micropsMWW89]
         |         Eviotops Smith 1956S58
         |           `--*E. infulatus Smith 1956S58
         |         Coryogalops Smith 1958S58
         |           `--*C. anomolus Smith 1958S58
         |         Mucogobius McCulloch 1912 [incl. Metagobius Whitley 1930]S58
         |           |--*M. mucosus (Gunther 1871) [=Gobius mucosus]S58
         |           |--M. bifasciatus Smith 1958S58
         |           |--M. flavobrunneus Smith 1958S58
         |           |--M. hasseltii Bleeker 1851S58
         |           |--M. insolita Whitley 1928S58
         |           `--M. liolepis (Koumans 1931)S58 [=Callogobius liolepisS58; incl. Macgregorella santa Herre 1931M58]
         |         Eleotriodes Bleeker 1858 (see below for synonymy)S58
         |           |--*E. sexguttatus (Valenciennes 1837)S58 (see below for synonymy)
         |           |--E. helsdingenii Bleeker 1858 [=Calleleotris helsdingenii]S58
         |           |--E. muralis Quoy & Gaimard in C & V 1837S58
         |           `--E. wardii Playfair 1866S58
         |         Cryptocentrops Smith 1958S58
         |           `--*C. exilis Smith 1958S58
         |         Pogonoculius Fowler 1938S58
         |           `--*P. zebra Fowler 1938S58
         |         Laccoeleotris Fowler 1935S58
         |           `--*L. lineopinnis Fowler 1935S58
         |         BatangaMWW89
         |         Krefftius Ogilby 1897O97
         |           `--*K. australis (Krefft 1864) [=Eleotris australis]O97
         |         Mulgoa Ogilby 1897O97
         |           `--*M. coxii (Krefft 1864) (see below for synonymy)O97
         |         Ophiorrhinus Ogilby 1897O97
         |           |--*O. grandiceps (Krefft 1864) [=Eleotris grandiceps]O97
         |           `--O. nudiceps (Castelnau 1873) [=Eleotris nudiceps, Philypnodon nudiceps]O97
         |         GymnobutisO97
         |           |--G. gymnocephalus (Steindachner 1867) [=Eleotris gymnocephalus]O97
         |           `--G. striatus Steindachner 1866O97
         |--+--Xenisthmus Snyder 1908T09, S58 [incl. Gignimentum Whitley 1933S58, Platycephalops Smith 1956S58; Xenisthmidae]
         |  |    |--*X. proriger Snyder 1908S58
         |  |    |--X. africanus Smith 1958S58
         |  |    |--X. clarus (Jordan & Seale 1905)MM09, S58 [=Hetereleotris claraS58]
         |  |    |--‘Gignimentum’ penicillum Whitley 1933S58
         |  |    `--X. polyzonatus (Klunzinger 1871) [=Eleotris polyzonatus]S58
         |  `--+--Ratsirakea legendreiT09
         |     |--Tateurndina ocellicauda Nichols 1955T09, M58
         |     `--+--OphieleotrisT09
         |        |    |--O. aporos (Bleeker 1854)T09, S58 (see below for synonymy)
         |        |    `--O. obscuraT90
         |        `--MogurndaT09
         |             |--M. adspersaT09
         |             |--M. australisG75
         |             |--M. bloodi Whitley 1956AC90a
         |             |--M. mogurnda (Richardson 1844) [=Eleotris mogurnda; incl. M. aurofodinae Whitley 1938]M58
         |             |--M. nesolepis (Weber 1908) [=Eleotris (Odonteleotris) nesolepis]AC90a
         |             |--M. oligolepis Allen & Jenkins 1999M10
         |             |--M. striataG75
         |             `--M. variegata Nichols 1951M58
            `--+--+--+--Microphilypnus ternetziT09
               |  |  `--LeptophilypnusT09
               |  |       |--L. fluviatilisT09
               |  |       `--L. panamensisT09
               |  `--+--GobiomorusT09
               |     |    |--G. maculatusT09
               |     |    `--+--G. dormitorT09
               |     |       `--Hemieleotris latifasciatusT09
               |     `--+--Guavina micropusT09
               |        `--DormitatorT09
               |             |--D. latifronsT09
               |             `--D. maculatusT09 [=Eleotris maculatusF16]
                  |  `--Calumia Smith 1958T09, S58
                  |       |--*C. biocellata Smith 1958S58
                  |       `--C. godfrayiT09
                     |    |--P. brevicepsS49
                     |    |--P. grandicepsT09
                     |    `--P. macrostomusT11
                     `--Gobiomorphus Gill 1863T09, H27
                          |  i. s.: G. basalisS49
                          |         G. cotidianusO81
                          |         G. huttoniH97
                          |         G. illotus Herre 1927H27
                          |--G. australisT09
                          `--+--G. coxiiT09
                             `--+--G. brevicepsT09
                                `--G. hubbsiT09
Eleotridae incertae sedis:
  Belobranchus Bleeker 1856H27
    `--B. belobranchus (Valenciennes 1837)M58 (see below for synonymy)
  Bunaka Herre 1927H27
    |--*B. pinguis Herre 1927H27
    |--B. gyrinoides (Bleeker 1853) [=Eleotris gyrinoides, Guavina gyrinoides]M58
    `--B. herwerdenii (Weber 1910) [=Eleotris herwerdenii, Oxyeleotris herwerdenii]M58
    |--F. cassandrae Nichols 1952M58
    `--F. diaphana Harry 1949M58
  Kimberleyeleotris Hoese & Allen 1987AC90b
    |--K. hutchinsiAL90
    `--K. notataAL90
  Grahamichthys radiataT11, H01
  Thalasseleotris [Thalasseleotrididae]T11
  Paloa Herre 1927H27
    `--*P. polylepis Herre 1927H27
  Parviparma Herre 1927H27
    `--*P. straminea Herre 1927H27
  Boroda Herre 1927H27
    |--*B. expatria Herre 1927H27
    `--B. albooculata Herre 1927H27

Belobranchus belobranchus (Valenciennes 1837)M58 [=Eleotris belobranchaM58; incl. B. quoyi Bleeker 1877H27, B. taeniopterus Bleeker 1856–1857H27, Eleotris taeniopteraH27]

Eleotriodes Bleeker 1858 [incl. Calleleotris Gill 1863, Gergobius Whitley 1930, Salarigobius Pfeffer 1893]S58

*Eleotriodes sexguttatus (Valenciennes 1837)S58 [=Eleotris sexguttatusS58, E. sexguttataS58; incl. E. lantzii Thominot 1878S58, Salarigobius stuhlmanni Pfeffer 1893S58, Valenciennea violifera MacCulloch & Ogilby 1919M58]

*Mulgoa coxii (Krefft 1864) [=Eleotris coxii; incl. E. mastersii Macleay 1881, E. richardsonii Steindachner 1866]O97

Ophieleotris aporos (Bleeker 1854)T09, S58 [=Eleotris aporosAC90a, Ophiocara aporosS58; incl. Ophiocara aporos var. guntheri Koumans 1937M58, Eleotris hoedtii Bleeker 1854M58, Ophiocara aporos hoedtiiM58, E. kuak Thiollière 1857M58, E. (Giurus) laglaizei Sauvage 1880H27, Ophiocara laglaiziJR10, Ophiocara aporos rigonis Whitley 1938M58]

*Type species of generic name indicated


[AC90a] Allen, G. R., & D. Coates. 1990a. An ichthyological survey of the Sepik River, Papua New Guinea. Records of the Western Australian Museum Supplement 34: 31–116.

[AC90b] Allen, G. R., & D. Coates. 1990b. A new species of freshwater eleotridid fish from northern Papua New Guinea. Records of the Western Australian Museum Supplement 34: 131–137.

[AL90] Allen, G. R., & R. Leggett. 1990. A collection of freshwater fishes from the Kimberley region of Western Australia. Records of the Western Australian Museum 14 (4): 527–545.

[F16] Fowler, H. W. 1916. The fishes of Trinidad, Grenada, and St. Lucia, British West Indies. Proceedings of the Academy of Natural Sciences of Philadelphia 67 (3): 520–548.

[G75] Grant, E. M. 1975. Guide to Fishes. The Co-ordinator-General’s Department: Brisbane (Australia).

[H01] Harcourt, R. G. 2001. Advances in New Zealand mammalogy 1990–2000: pinnipeds. Journal of the Royal Society of New Zealand 31 (1): 135–160.

[H27] Herre, A. W. 1927. Gobies of the Philippines and the China Sea. Bureau of Printing: Manila.

[H97] Hicks, B. J. 1997. Food webs in forest and pasture streams in the Waikato region, New Zealand: a study based on analyses of stable isotopes of carbon and nitrogen, and fish gut contents. New Zealand Journal of Freshwater and Marine Research 31: 651–664.

[MWW89] Miller, P. J., J. Wright & P. Wongrat. 1989. An Indo-Pacific goby (Teleostei: Gobioidei) from West Africa, with systematic notes on Butis and related eleotridine genera. Journal of Natural History 23: 311–324.

[MM09] Moore, G., & S. Morrison. 2009. Fishes of three North West Shelf atolls of Western Australia: Mermaid (Rowley Shoals), Scott and Seringapatam Reefs. Records of the Western Australian Museum Supplement 77: 221–255.

[M10] Morgan, D. L. 2010. Fishes of the King Edward River in the Kimberley region, Western Australia. Records of the Western Australian Museum 25 (4): 351–368.

[M58] Munro, I. S. R. 1958. The fishes of the New Guinea region: a check-list of the fishes of New Guinea incorporating records of species collected by the Fisheries Survey Vessel “Fairwind” during the years 1948 to 1950. Papua and New Guinea Agricultural Journal 10 (4): 97–369 (reprinted: 1958. Territory of Papua and New Guinea Fisheries Bulletin no. 1).

[O81] O’Brien, C. O. 1981. A. A. Book of New Zealand Wildlife: A guide to the native and introduced animals of New Zealand. Lansdowne Press: Auckland.

[O97] Ogilby, J. D. 1897. On some Australian Eleotrinae. Proceedings of the Linnean Society of New South Wales 21 (4): 725–757.

[S58] Smith, J. L. B. 1958. The fishes of the family Eleotridae in the western Indian Ocean. Ichthyological Bulletin 11: 137–163.

[S49] Stokell, G. 1949. The systematic arrangement of the New Zealand Galaxiidae. Part II. Specific classification. Transactions and Proceedings of the Royal Society of New Zealand 77 (4): 472–496.

[T90] Taguchi, S. 1990. Nihon no Sakana. Kogakukan: Tokyo.

[T09] Thacker, C. E. 2009. Phylogeny of Gobioidei and placement within Acanthomorpha, with a new classification and investigation of diversification and character evolution. Copeia 2009 (1): 93–104.

[T11] Thacker, C. 2011. Systematics of Butidae and Eleotridae. In: Patzner, R. A., J. L. Van Tassell, M. Kovačić and B. G. Kapoor (eds) The Biology of Gobies pp. 79–85. CRC Press.

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