Northern horsebalm Collinsonia canadensis, copyright Michael Hough.

Belongs within: Lamiaceae.

The Elsholtzieae are a group of herbs and shrubs with an asymmetric floral disc and globose nutlets (Harley et al. 2004).

Characters (from Harley et al. 2004): Annual or perennial herbs or shrubs. Stems often four-angled. Leaves opposite, usually petiolate. Inflorescence usually an often secund raceme-like thyrse or panicle of 1(-3)-flowered cymes in the axils of usually much reduced bracts, sometimes conspicuous, broad, imbricate, often venose, bracteoles usually absent. Calyx ± 10-lobed, five-nerved (3/2), often more or less actinomorphic at anthesis, usually accrescent in fruit and usually becoming more distinctly two-lipped, corolla ± 2-lipped, 4-5-lobed (1-2/3), lobes usually with short, scarcely spreading lobes, posterior lip 1-2-lobed, anterior lip 3-lobed, corolla-tube often short. Stamens 4 or 2 by reduction, often exserted, weakly didynamous, divergent, filaments glabrous, anthers 2-thecous, thecae parallel to divergent, separate or apically confluent. Style with two usually equal, subulate stigma-lobes. Disc asymmetrical with anterior lobe elongate. Nutlets usually globose, sometimes reduced to one by abortion, smooth, reticulate or foveolate, abscission-scar very small, often circular.

    |    |--E. densaO88
    |    |--E. eriostachyaO88
    |    |--E. flavaSS72
    |    |--E. fruticosaO88
    |    |--E. stachyodiaSS72
    |    `--E. strobiliferaO88
    `--Collinsonia Linnaeus 1753 [incl. Keiskea Miquel 1865]HPR03
         |--*C. canadensisHPR03
         |--C. australis (Wu & Li) Harley in Harley, Paton & Ryding 2003 [=Keiskea australis Wu & Li 1977]HPR03
         |--C. elsholtzioides (Merr.) Harley in Harley, Paton & Ryding 2003 [=Keiskea elsholtzioides Merr. 1937]HPR03
         |--C. glandulosa (Wu) Harley in Harley, Paton & Ryding 2003 [=Keiskea glandulosa Wu 1965]HPR03
         |--C. japonica (Miquel) Harley in Harley, Paton & Ryding 2003 [=*Keiskea japonica Miquel 1865]HPR03
         |--C. macrobracteata (Masam.) Harley in Harley, Paton & Ryding 2003 [=Keiskea macrobracteata Masam. 1940]HPR03
         |--C. sinensis (Diels) Harley in Harley, Paton & Ryding 2003 [=Keiskea sinensis Diels 1924]HPR03
         `--C. szechuanensis (Wu) Harley in Harley, Paton & Ryding 2003 [=Keiskea szechuanensis Wu 1965]HPR03

*Type species of generic name indicated


[HPR03] Harley, R. M., A. J. Paton & O. Ryding. 2003. New synonymy and taxonomic changes in the Labiatae. Kew Bulletin 58: 485–489.

[O88] Ohba, H. 1988. The alpine flora of the Nepal Himalayas: an introductory note. In: Ohba, H., & S. B. Malla (eds) The Himalayan Plants vol. 1. The University Museum, University of Tokyo, Bulletin 31: 19–46.

[SS72] Singh, V., & H. Singh. 1972. A contribution to the flora of Gangolihat block in Pithoragarh District. Journal of the Bombay Natural History Society 69 (2): 352–368.

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