Empidoidea

Protempis antennata, from Grimaldi & Engel (2005).

Belongs within: Neoneura.
Contains: Hybotidae, Dolichopodidae, Empididae.

The Empidoidea are a clade of long-legged flies supported by molecular data. Members of this clade are predators as adults and larvae (Grimaldi & Engel 2005). In the past this group has been divided between the families Dolichopodidae and Empididae but recognition of the latter as paraphyletic has led to its division between the Empididae sensu stricto, Atelestidae, Hybotidae and basal Dolichopodidae. The Atelestidae have the costa ending at or near vein M1+2, vein R5 originating as far or further from the humeral crossvein than the length of said crossvein, the prosternum and proepisternum separated, and symmetrical, unrotated male terminalia (Wahlberg & Johanson 2018).

<==Empidoidea [Empidi, Oreogetoninae]
    |  i. s.: GondwanamyiaWJ18
    |--Protoreogeton admirabilisGE05
    |--Homalocnemimus abjugatusGE05
    |--Protempis [Protempididae, Protempidinae]GE05
    |    |--P. antennataGE05
    |    `--P. minuta Ran 1998GC99
    `--+--+--HybotidaeWJ18
       |  `--+--DolichopodidaeWJ18
       |     `--+--EmpididaeWJ18
       |        `--RagadidaeWJ18
       |             |--Iteaphila Zetterstedt 1840B89 [IteaphilinaeWJ18]
       |             |    |  i. s.: I. coriacea Bigot 1889B89
       |             |    |--I. macquartiSK81
       |             |    `--+--I. furcataWJ18
       |             |       `--AnthepiscopusSK81
       |             `--RagadinaeWJ18
       |                  |--DipsomyiaWJ18
       |                  |--Hormopeza Zetterstedt 1840WJ18, B89
       |                  |    `--H. brevicornisSK81
       |                  |--HydropezaWJ18
       |                  |--ZanclotusWJ18
       |                  `--Ragas Walker 1837B89
       |                       |--R. primigeniaSK81
       |                       `--R. unicaWJ18
       `--AtelestidaeWT11
            |  i. s.: Dianafranskia fisheriGE05
            |--AtelestinaeWJ18
            |    |--AcarteropteraWJ18
            |    |--Alavesia subiasiWJ18, GE05
            |    |--MeghyperusWJ18
            |    |--AtelestusWT11 [incl. PlatycnemaK87]
            |    |     `--A. pulicariusWT11
            |    `--Atelestites Grimaldi & Cumming 1999GC99
            |         `--*A. senectus Grimaldi & Cumming 1999GC99
            `--+--Burmitempis halteralis Cockerell 1917GC99
               |--Nemedina [Nemedininae]WJ18
               |    `--N. alamirabilisGC99
               |--Prolatomyia Grimaldi & Cumming 1999GC99
               |    `--*P. elongata Grimaldi & Cumming 1999GC99
               |--Cretodromia Grimaldi & Cumming 1999GC99
               |    `--*C. glaesa Grimaldi & Cumming 1999GC99
               |--Phaetempis Grimaldi & Cumming 1999GC99
               |    `--*P. lebanensis Grimaldi & Cumming 1999GC99
               |--Nemedromia Grimaldi & Cumming 1999GC99
               |    |--*N. campania Grimaldi & Cumming 1999GC99
               |    |--N. telescopica Grimaldi & Cumming 1999GC99
               |    `--N. turonia Grimaldi & Cumming 1999GC99
               `--Neoturonius Grimaldi & Cumming 1999GC99
                    |--*N. asymmetrus Grimaldi & Cumming 1999GC99
                    |--N. cretatus Grimaldi & Cumming 1999GC99
                    `--N. vetus Grimaldi & Cumming 1999GC99

*Type species of generic name indicated

References

[B89] Bigot, J.-M.-F. 1889. Diptères nouveaux ou peu connus. 34e partie. XLII. Empidi. Annales de la Société Entomologique de France, 6e série 9: 111–134.

[GC99] Grimaldi, D., & J. Cumming. 1999. Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura. Bulletin of the American Museum of Natural History 239: 1–124.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[K87] Kessel, E. L. 1987. Platypezidae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 681–688. Research Branch, Agriculture Canada.

[SK81] Steyskal, G. C., & L. V. Knutson. 1981. Empididae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 607–624. Research Branch, Agriculture Canada.

[WJ18] Wahlberg, E., & K. A. Johanson. 2018. Molecular phylogenetics reveals novel relationships within Empidoidea (Diptera). Systematic Entomology 43: 619–636.

[WT11] Wiegmann, B. M., M. D. Trautwein, I. S. Winkler, N. B. Barr, J.-W. Kim, C. Lambkin, M. A. Bertone, B. K. Cassel, K. M. Bayless, A. M. Heimberg, B. M. Wheeler, K. J. Peterson, T. Pape, B. J. Sinclair, J. H. Skevington, V. Blagoderov, J. Caravas, S. N. Kutty, U. Schmidt-Ott, G. E. Kampmeier, F. C. Thompson, D. A. Grimaldi, A. T. Beckenbach, G. W. Courtney, M. Friedrich, R. Meier & D. K. Yeates. 2011. Episodic radiations in the fly tree of life. Proceedings of the National Academy of Sciences of the USA 108 (14): 5690–5695.

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