Endothyrinae

Endothyra baileyi, from Lousville Fossils.

Belongs within: Endothyroida.

The Endothyrinae are a group of enrolled foraminifera known from the Upper Devonian to the Permian. The plane of coiling of the test often changes during growth, sometimes gradually (plectogyral growth) or sometimes with an abrupt turn of up to 90°. Members of the genera Endothyranella and Rectoseptaglomospiranella become uncoiled and rectilinear in later sections of the test (Loeblich & Tappan 1964a).

Characters (from Loeblich & Tappan 1964a): Test enrolled, planispiral to streptospiral, possibly uncoiling in later stages; may be incompletely septate; wall of granular calcite, nonporous, with two or three distinct layers in well-preserved specimens, may have agglutinated material in addition; aperture single, simple, basal or areal.

<==Endothyrinae [Plectogyrinae]
    |--+--Septammina Meunier 1888C40
    |  |    |--*S. renaulti Meunier 1888C40
    |  |    `--S. bradyiC40
    |  `--Endothyranella Galloway & Harlton 1930C40, R-CB96
    |       |--*E. powersi (Harlton 1927) [=Ammobaculites powersi]LT64a
    |       `--E. gracilisR-CB96
    `--Endothyra Phillips 1846 (nom. cons.)C40 [incl. Plectogyra Zeller 1950LT64a]
         |  i. s.: E. baileyiLT64a
         |         E. crassaG31
         |         E. mediaC40
         |         E. plectogyra (Zeller 1950) [=*Plectogyra plectogyra]LT64a
         |         E. timanica Durkina 1959CV01
         |--E. (Endothyra)R-CB96
         |    |--*E. (E.) bowmani Phillips 1846 (nom. cons.)LT64a (see below for synonymy)
         |    `--E. (E.) phrissaR-CB96
         |--E. (Latiendothyra Lipina 1963)R-CB96
         |    `--E. (*L.) latispiralis Lipina 1955R-CB96
         |--E. (Mediendothyra Brazhnikova & Vdovenko 1972)R-CB96
         |    `--E. (*M.) obscura Brazhnikova & Vdovenko 1971R-CB96
         |--E. (Plectogyrina Reitlinger 1959)R-CB96
         |    `--E. (*P.) fomichaensis Lebedeva 1954LT64a
         |--E. (Similisella Vdovenko in Rauzer-Chernousova, Bensh et al. 1996)R-CB96
         |    `--E. (*S.) similis Rauser & Reitlinger 1936R-CB96
         |--E. (Spirella Vdovenko in Rauzer-Chernousova, Bensh et al. 1996)R-CB96
         |    `--E. (*S.) spira (Conil & Lys 1964) [=Plectogyra spira]R-CB96
         `--E. (Tuberendothyra Skipp 1969)R-CB96
              `--E. (*T.) tuberculata Lipina 1948R-CB96
Endothyrinae incertae sedis:
  Pseudonovella Kireeva 1949LT64b
  Melatolla Strank 1983R-CB96
    `--*M. whitfieldensis Strank 1983R-CB96
  Mikhailovella Ganelina 1956R-CB96
    `--*M. gracilis (Rauser 1948)R-CB96 (see below for synonymy)
  Planoendothyra Reitlinger 1959R-CB96
    |--*P. (Planoendothyra) aljutovica (Reitlinger 1950)R-CB96 (see below for synonymy)
    `--P. (Iriclinella Malakhova 1980)R-CB96
         `--P. (*I.) evoluta (Reitlinger 1950) [=Endothyra spirilliniformis var. evoluta]R-CB96
  Priscella Mamet 1974R-CB96
    `--*P. prisca (Rauser & Reitlinger 1936) [=Endothyra prisca]R-CB96
  Spinothyra Mamet 1976R-CB96
    `--*S. pauciseptata (Rauser 1948) [=Endothyra pauciseptata]R-CB96
  Rectoendothyra Brazhnikova 1983R-CB96
    `--*R. donbassica Brazhnikova 1983R-CB96

*Endothyra (Endothyra) bowmani Phillips 1846 (nom. cons.)LT64a [incl. E. bradyi Mikhailov 1939R-CB96, Involutina lobata Brady 1870LT64a]

*Mikhailovella gracilis (Rauser 1948)R-CB96 [=Endothyrina gracilisLT64a, Haplophragmella gracilisLT64a, Melatolla gracilisR-CB96]

*Planoendothyra (Planoendothyra) aljutovica (Reitlinger 1950)R-CB96 [=Endothyra aljutovicaR-CB96, Quasiendothyra aljutovicaLT64a]

*Type species of generic name indicated

References

[CV01] Cózar, P., & D. Vachard. 2001. Dainellinae subfam. nov. (Foraminiferida du Carbomifère inférieur), révision et nouveaux taxons. Geobios 34 (5): 505–526.

[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3rd ed. Harvard University Press: Cambridge (Massachusetts).

[G31] Grabau, A. W. 1931. The Permian of Mongolia: A report on the Permian fauna of the Jisu Honguer limestone of Mongolia and its relations to the Permian of other parts of the world. American Museum of Natural History: New York.

[LT64a] Loeblich, A. R., Jr & H. Tappan. 1964a. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[LT64b] Loeblich, A. R., Jr & H. Tappan. 1964b. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 2. The Geological Society of America and The University of Kansas Press.

[R-CB96] Rauzer-Chernousova, D. M., F. R. Bensh, M. V. Vdovenko, N. B. Gibshman, E. Y. Leven, O. A. Lipina, E. A. Reitlinger, M. N. Solovieva & I. O. Chedija. 1996. Spravočnik po Sistematike Foraminifer Paleozoâ (Èndotiroidy, Fuzulinoidy). Rossijskaâ Akademiâ Nauk, Geologičeskij Institut, Moskva “Nauka”.

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