Engina alveolata, copyright G. & Ph. Poppe.

Belongs within: Pisaniidae.

Engina is a genus of tropical marine gastropods with a sharp-spired, ovate shell bearing prominent nodulous ribs.

Characters (from Oldroyd 1927): Shell ovate-conic; spire shapr, with longitudinal nodulous ribs, decussated by revolving lines; aperture narrow, with several oblique plications in front; outer lip rather thick, inflected or callous in middle, crenulated within.

Engina Gray 1839 [incl. Pusiostoma Swainson 1840; Engininae, Pusiostomatidae]BR05
|--E. turbinella (Kiener 1836)KF22 (see below for synonymy)
`--+--E. alveolata (Kiener 1836)KF22, F27 (see below for synonymy)
`--E. mendicaria (Linnaeus 1758)KF22, BR17 [=Voluta mendicariaH09, *Pusiostoma mendicariaBR17]

Engina incertae sedis:
E. albocincta Pease 1860 [=Columbella (Engina) albocincta]K65
E. anaxeres (Kiener 1836)H09 [=Purpura anaxeresH09, Sistrum anaxaresM01]
E. armillata (Reeve 1846)W94
E. australis (Pease 1871) [=Maculotriton australis, Tritonidea australis]W94
E. bonasia (von Martens 1880)W94 [=Plicatella (Peristernia) bonasiaC80a; incl. E. zatricum Melvill 1893W94]
E. carbonaria Reeve 1846O27
E. carduus (Reeve 1844) [=Triton carduus, Nassaria carduus]C80b
E. concinna (Reeve 1846)H09 [=Ricinula concinnaH09; incl. Tritonidea submenkeana Pilsbry 1901C80b, W94]
E. costata Pease 1860 [=Columbella (Engina) costata]K65
E. crocostomaC64
E. cronuchorda Fraussen & Chino 2011WBS-S15
E. curtisiana (Smith 1884) [incl. E. sinensis Melvill 1895]W94
E. farinosaC80b
E. ‘fusiformis’ Stearns 1894 non Pease 1865PP78
E. incarnata (Deshayes 1834) (see below for synonymy)W94
E. lineata (Reeve 1846)H09 [=Ricinula lineataH09; incl. E. lineata var. maculata Pease 1869W94]
E. menkeana (Dunker 1860) [=Cantharus (Pollia) menkeana, C. menkeanus]C80b
E. monilifera Pease 1860 (see below for synonymy)K65
E. ovata Pease 1865K65
E. phasinola (Duclos 1840)W94 [=Columbella phasinolaC80a]
E. philippiii (Cossmann 1899) [=Sveltella philippii, Cancellaria parva Philippi 1860 non Lea 1833]PH90
E. recurvaB79
E. resta (Iredale 1940)W94 [=Enzinopsis restaC80b]
E. ryalli Rolán & Fernandes 1995BC01
E. siderea (Reeve 1846) [=Ricinula siderea]H09
E. trifasciata (Reeve 1846) [=Ricinula trifasciata]H09
E. tuberculosa Pease 1863K65
E. zonalis (Lamarck 1822) [incl. E. melanozona Tomlin 1928]W94

Engina alveolata (Kiener 1836)KF22, F27 [=Purpura alveolataF27; incl. E. fusiformis Pease 1865W94, Columbella (Engina) fusiformis nec Colombella fusiformis Anton 1839 nec Colombella fusiformis d’Orbigny in Sagra 1842 (not preoc. if in dif. gen.)K65, E. histrio Reeve 1846W94, Ricinula lauta Reeve 1846H09, W94, E. lautaH09, E. mundula Melvill & Standen 1895W94, E. reevei Tryon 1883W94]

Engina incarnata (Deshayes 1834) [incl. E. astricta Reeve 1846, E. kobeltiana Tapparone-Canefri 1880, E. paulucciae Tapparone-Canefri 1879]W94

Engina monilifera Pease 1860 [=Columbella (Engina) monilifera nec Columbella monilifera Sowerby 1844 nec Colombella monilifera Duclos in Chenu 1846 (not preoc. if in dif. gen.), Enzina (l. c.) monilifera]K65

Engina turbinella (Kiener 1836)KF22 [=Purpura turbinellaKF22; incl. E. tubinella var. cruzana Usticke 1959BC01, *E. zonata Gray 1839KF22]

*Type species of generic name indicated


[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[B79] Brazier, J. 1879. List of marine shells collected on Fitzroy Island, north coast of Australia; with notes on their geographical range. Journal of Conchology 2: 186–199.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[C80a] Cernohorsky, W. O. 1980a. The taxonomy of some Indo-Pacific Mollusca. Part 7. Records of the Auckland Institute and Museum 16: 171–187.

[C80b] Cernohorsky, W. O. 1980b. The taxonomy of some Indo-Pacific Mollusca. Part 8. Records of the Auckland Institute and Museum 17: 135–152.

[F27] Finlay, H. J. 1927. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[KF22] Kantor, Y. I., A. E. Fedosov, A. R. Kosyan, N. Puillandre, P. A. Sorokin, Y. Kano, R. Clark & P. Bouchet. 2022. Molecular phylogeny and revised classification of the Buccinoidea (Neogastropoda). Zoological Journal of the Linnean Society 194 (3): 789–857.

[K65] Kay, E. A. 1965. Marine molluscs in the Cuming collection, British Museum (Natural History) described by William Harper Pease. Bulletin of the British Museum (Natural History): Zoology Supplement 1: 1–96, 14 pls.

[M01] Melville, J. C. 1901. A few further remarks upon the Erythrean molluscan fauna, with descriptions of seven species from Aden, in the collection of Commander E. R. Shopland, R.I.M. Annals and Magazine of Natural History, series 7, 7: 550–556.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. 2 pt 1. Stanford University Press: Stanford University (California).

[PH90] Petit, R. E., & M. G. Harasewych. 1990. Catalogue of the superfamily Cancellarioidea Forbes and Hanley, 1851 (Gastropoda: Prosobranchia). Nautilus 103 (Suppl. 1): 1–69.

[PP78] Poorman, F. L., & L. H. Poorman. 1978. Additional molluscan records from Bahía de Los Angeles, Baja California Norte. Veliger 20 (4): 369–374.

[WBS-S15] Willan, R. C., C. Bryce & S. M. Slack-Smith. 2015. Kimberley marine biota. Historical data: molluscs. Records of the Western Australian Museum Supplement 84: 287–343.

[W94] Wilson, B. 1994. Australian Marine Shells vol. 2. Prosobranch Gastropods. Part Two (Neogastropods). Odyssey Publishing: Kallaroo (Western Australia).

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