Entiminae

Lachnopus floridanus, copyright David Victoria.

Belongs within: Cyclominae.
Contains: Peritelini, Sciaphilini, Pachyrhynchini, Eupholini, Alophini, Polydrusini, Brachyderini, Cyphicerini, Otiorhynchini, Tropiphorini, Trachyphloeini, Phyllobiini, Tanymecini, Eustylini, Geonemini, Leptopiini, Naupactini.

The Entiminae, broad-nosed weevils, have a shorter, broader snout than most other weevils. They typically bear a deciduous process on the mandibles that breaks off shortly after emergence, leaving a distinct scar. The apex of the hind tibia bears only a short tooth or spine on the inner angle. Many species are flightless. Larvae are typically external feeders on roots in soil. Both adults and larvae are commonly polyphagous (Anderson 2002).

<==Entiminae (see below for synonymy)
|--+--+--+--PolydrusiniLE23
| | | `--Laparocerus [Laparocerini]GC-P14
| | | `--L. freyiGC-P14
| | `--+--Barianus [Oosomini]GC-P14
| | `--BrachyderiniGC-P14
| `--+--CyphiceriniA02
| `--+--+--+--OtiorhynchiniHL23
| | | `--TropiphoriniGC-P14
| | `--+--MyorhinusLC40 [MyorhininiGC-P14]
| | | `--M. albolineatus [=Curculio albolineatus]LC40
| | `--TrachyphloeiniGC-P14
| `--PhyllobiiniHL23
`--+--+--+--+--Cratopus [Cratopini]GC-P14
| | | | `--C. marmoreusLC40
| | | |--TanymeciniGC-P14
| | | `--EustyliniPLB13
| | `--+--GeoneminiHL23
| | `--Psallidium [Psallidiini]GC-P14
| | |--P. desbrochersi Faust 1894HF03
| | |--P. maxillosumC01
| | `--P. separandum Desbrochers 1875HF03
| `--+--OphryastiniGC-P14
| | |--Ophryastes Germar 1829 (see below for synonymy)A02
| | | |--O. sulcirostrisBM76
| | | `--O. vittatus [=Liparus vittatus]LC40
| | `--Sapotes Casey 1888A02
| | |--S. longipilis Van Dyke 1934A02
| | `--S. puncticollis Casey 1888A02
| `--Lachnopus Schoenherr 1840GC-P14, FS90 [incl. Menoetius Dejean 1821A02, Ptilopus Schoenherr 1823A02]
| |--L. argus (Reiche 1840)A02
| |--‘Curculio’ aurifer [=Ptilopus aurifer]LC40
| |--L. campechianus Chv. 1840 [incl. L. adspersus Chv. 1880]FS90
| |--L. curvipes Fab. 1787FS90
| |--L. floridanus Horn 1876A02
| |--L. hispidus (Gyllenhal 1834)A02
| `--L. lineicollis Chv. 1880 [incl. L. foveicollis Chv. 1880, L. quadritaenia Chv. 1880]FS90
`--+--LeptopiiniHL23
`--+--EntimusHL23 [EntiminiPLB13]
| |--E. imperialiaHL23
| |--E. nobilisLC40
| `--E. splendidus [=Curculio splendidus]LC40
`--NaupactiniHL23

Entiminae incertae sedis:
Achradidus milleri (Tournier 1874)HF03
Conocoetus baudii (Faust 1889)HF03
ParascythopusHF03
StrophomorphusHF03
|--S. colonnellii Pelletier 1999HF03
|--S. impressicollis Tournier 1874HF03
|--S. milleri Tournier 1874HF03
|--S. minutus Tournier 1874HF03
|--S. perrinae Pelletier 1999HF03
`--S. ventricosus (Chevrolat 1880)HF03
Dorotheus guidensis Kuschel 1959PLB13
Arostropsis groehni Yunakov & Kirejtshuk 2011PLB13
PaonaupactusPLB13
|--P. cephalotes (Voss 1972)PLB13
`--P. sitonitoides Voss 1953PLB13
ProtonaupactusPLB13
|--P. microphthalmus Zherikhin 1971PLB13
|--P. sobrinus (Voss 1972)PLB13
`--P. viridis (Wanat & Borowiec 1986)PLB13
Archaeoscipahilus marshalli Legalov 2012PLB13
Archaeocallirhopalus larssoni Legalov 2013PLB13
PromecopsPLB13
|--P. episcopalisLC40
|--P. postica Fåhrs. 1840 [incl. P. alboscutellata Chv. 1880, P. lateralis]FS90
`--P. tumidirostris Poinar & Brown 2011PLB13
Gymnopholus (Symbiopholus) lichenifer Gressitt 1966B14
Apirocalus cornutusMac86
CoptorhynchusMas86
|--C. bombicollis Macleay 1886Mac86
|--C. gestroi Pascoe 1885Mas86
|--C. jansoni Pascoe 1871Mas86
|--C. lateralisMas86
|--C. melancholicusMas86
|--C. nudus Macleay 1886Mac86
|--C. scabratusMas86
|--C. speculatus Macleay 1886Mac86
`--C. tessellatusMac86
Celeuthetes echinatus (Fabricius 1801)Z93 [=Curculio echinatusZ93; incl. Ce. australisMas86, Ce. paganusMas86]
Psomeles oblongusMas86
TyloderesMas86
|--T. chrysops [=Curculio chrysops]LC40
`--T. spinosus Boisduval 1835Mas86
MerimnetesMas86
|--M. tenuis Germ. 1848Mas86
`--M. uniformisMas86
Euphalia pardalis Pascoe 1870Mas86
ProxyrusMas86
|--P. abstersus Pascoe 1870Mas86
`--P. lecideosus Pascoe 1870Mas86
AtmesiaMas86
|--A. glaucina Pascoe 1873Mas86
`--A. marginata Pascoe 1870Mas86
TimaretaMas86
|--T. crinita Pascoe 1874Mas86
|--T. figurata Pascoe 1872Mas86
`--T. satellina Pascoe 1872Mas86
Phylda periteloides Pascoe 1869Mas86
TitiniaMas86
|--T. ignaria Pascoe 1869Mas86
`--T. marmorata Pascoe 1872Mas86
Idaspora terrea Pascoe 1869Mas86
TelenicaMas86
|--T. nebulosa Pascoe 1872Mas86
`--T. sublimbata Pascoe 1872Mas86
Epherina longicornis Pascoe 1869Mas86
Synomus cephalotes Pascoe 1885Mas86
Bothynorrhynchus [incl. Opetiopteryx]Z93
`--B. lascivusMas86
Psapharus Schoenherr 1823 [incl. Hybauchinia]Z93
|--*P. infaustusZ93 [=Curculio infaustusZ93, Cherrus infaustusB35; incl. Ch. bulbiferMas86, Ch. granulatusMas86]
`--P. mistothes [incl. Hybauchinia nodulosa]Z93
DeracanthusZ93
|--*D. spinifex [=Curculio spinifex]Z93
`--D. pallasiiC01
Stereogastrus scabratus (Fabricius 1801) [=Curculio scabratus]Z93
Asceparnus nodipennis Schoenherr 1847Z93
Platyapistes argentinensisML05
Rhinoscaphus doriae Pascoe 1885Mac86
Apocyrtus froggatti Macleay 1886Mac86
Eutinophoea nana Pascoe 1870Mas86
Euthyphasis acuta Pascoe 1875Mas86
Acalanoma reducta Pascoe 1875Mas86
Ochrometa amoena Pascoe 1870Mas86
DermatodesMas86
|--D. australis Boisd. 1835Mas86
|--D. caesicollisF89
`--D. truncatirostris Fairmaire 1889F89
Laodice funebris Chev. 1880Mas86
ProstomusMas86
|--P. australis Boisduval 1835B35
|--P. scutellaris (Fabricius 1787) [=Curculio scutellaris; incl. C. exsertus Fabricius 1801]Z93
|--P. squalidusB35
`--P. tenebricosusB35
Styreus geonomoides Pascoe 1883Mas86
EvasMas86
|--E. acuminata Pascoe 1870Mas86
|--E. argenteiventris Pascoe 1870Mas86
|--E. crassirostris Pascoe 1870Mas86
`--E. lineatus Pascoe 1873Mas86
ProsauliusMas86
|--P. ateropterusMas86
|--P. comosus Germ. 1848Mas86
|--P. dispar Germ. 1848Mas86
`--P. hopeiMas86
PrypnusMas86
|--P. canaliculatusMas86
|--P. fallaxMas86
|--P. quinquenodosusMas86
|--P. squalidus [incl. P. bidentatus]Mas86
|--P. subtuberculatusMas86
|--P. tenebricosusMas86
`--P. trituberculatus Germ. 1848Mas86
AnypotactiniPLB13
|--Anypotactus Schoenherr 1840PLB13
`--Polydacrys Schoenherr 1834A02
|--P. depressifrons Boheman 1840A02
`--P. modestus Gyll. 1834 (see below for synonymy)FS90
CylydrorhinusW41 [CylydrorhininiPLB13]
`--C. angulatus Guerin-Meneville 1841W41
EudiagoginiPLB13
|--Eudiagogus Schoenherr 1840A02
|--Aracanthus Say 1831A02
| `--A. pallidus Say 1831A02
`--Colecerus Schoenherr 1840 [=Coleocerus Agassiz 1846; incl. Bathyris LeConte 1874]A02
|--C. dispar (LeConte 1874)A02
`--C. marmoratus (Horn 1876)A02
HormoriniPLB13
|--Agasphaerops Horn 1876A02
| `--A. niger Horn 1876A02
|--Hormorus Horn 1876A02
| `--H. undulatus (Uhler 1856)A02
`--Lupinocolus Van Dyke 1936A02
`--L. blaisdelli Van Dyke 1936A02
LordopsLC40 [LordopiniPLB13]
`--L. gyllenhali [=Curculio gyllenhali, Hypsonotus gyllenhali; incl. H. dives]LC40
CneorhininiPLB13
|--Philopedon Schoenherr 1826 (see below for synonymy)A02
| `--P. plagiatum (Schaller 1783)A02
`--CneorhinusB35
|--C. geminatus [=Curculio geminatus, Thylacites geminatus; incl. Cu. globatus]LC40
|--C. impressipennis Boisduval 1835B35
`--C. stygmatipennis Boisduval 1835B35
PeriteliniPLB13
SciaphiliniPLB13
OttistiriniM94
|--Ottistira fasciata Macleay 1886Mac86
`--Maleuterpes Blackburn 1893M94
|--*M. spinipes Blackburn 1893M94
`--M. phytolymusB70
PachyrhynchiniM94
EupholiniB14
AgraphiniA02
|--Agraphus Say 1831A02
| |--A. bellicus (Say 1831)A02
| `--A. leucophaeusLC40
`--Paragraphus Blatchley 1916A02
`--P. setosus Blatchley 1916A02
AlophiniA02
Omias Germar 1817 [incl. Mylacus Boheman 1843, Omorus Gistel 1856; Omiini]A02
|--O. indutusDL90
|--‘Mylacus’ rotundatusC01
`--O. seminolum [=Curculio seminolum]LC40
EpisomusLB20
|--E. chlorostigmaLB20
|--E. pauperatusP66
`--E. platina [=Curculio platina]LC40
OribiusLB20
NothesLB20
AmycterodontusZ93
Cyanippeus angulatusLC40
Arhines languidusLC40
Amblyrhinus poricollisLC40
Piezonotus suturalisLC40
Hadrorhinus lepidopterusLC40
PtochusLC40
|--P. circumcinctusLC40
`--P. deportatusC01
Cosmorhinus cristatusLC40
Pholicodes belbejusLC40
Liparocerus morioLC40
Anomus pubescens [=Aomus pubescens]LC40
Sciobius pullus [=Curculio pullus]LC40
Sphaeromus australisLC40
Hyphantus bacciferLC40
Elytrodon bidentatus [=Elytrodes bidentatus]LC40
Chloebius immeritusLC40

Entiminae [Brachyderides, Brachyderinae, Entimides, Leptopiinae, Otiorhynchides, Otiorrhynchinae, Polydrosinae, Thylacitinae]

Ophryastes Germar 1829 [incl. Amydrogmus Pierce 1913, Caccophryastes Sharp 1891, Dystirus Pascoe 1872, Eupagoderes Horn 1876, Tosastes Sharp 1891]A02

Philopedon Schoenherr 1826 [=Philopedum Agassiz 1846; incl. Dactylorhinus Tournier 1876, Dactylorrhinus Rye 1878]A02

Polydacrys modestus Gyll. 1834 [incl. P. luctuosus, P. modestus var. moestus Chv. 1880, P. modestus var. nigrosparsus Chv. 1880, P. scansorius]FS90

*Type species of generic name indicated

References

[A02] Anderson, R. S. 2002. Curculionidae Latreille 1802. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 722–815. CRC Press.

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[B35] Boisduval, J. B. 1835. Voyage de Découvertes de l’Astrolabe. Exécuté par ordre du Roi, pendant les années 1826–1827–1828–1829, sous le commandement de M. J. Dumont d’Urville. Faune entomologique de l’océan Pacifique, avec l’illustration des insectes nouveaux recueillis pendant le voyage vol. 2. Coléoptères et autres ordres. J. Tastu: Paris.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[B70] Britton, E. B. 1970. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 495–621. Melbourne University Press.

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[DL90] Desbrochers des Loges, J. 1890. Quelques observations au sujet de la synonymie de diverses espèces du genre Scythropus et de plusieurs autres espèces du même groupe. Annales de la Société Entomologique de France, 6e série 9: ccxxv–ccxxvi.

[F89] Fairmaire, L. 1889. Coléoptères de l’intérieur de la Chine. 5e partie. Annales de la Société Entomologique de France, 6e série 9: 5–84.

[FS90] Fleutiaux, E., & A. Sallé. 1890. Liste des coléoptères de la Guadeloupe et descriptions d’espèces nouvelles. Annales de la Société Entomologique de France, 6e série 9: 351–484.

[GC-P14] Gillett, C. P. D. T., A. Crampton-Platt, M. J. T. N. Timmermans, B. H. Jordal, B. C. Emerson & A. P. Vogler. 2014. Bulk de novo mitogenome assembly from pooled total DNA elucidates the phylogeny of weevils (Coleoptera: Curculionoidea). Molecular Biology and Evolution 31 (8): 2223–2237.

[HF03] Halperin, J., & J. Fremuth. 2003. Contribution to the knowledge of Curculionoidea (Coleoptera) and their host plants in Israel. Zoology in the Middle East 29: 93–100.

[HL23] Haran, J., X. Li, R. Allio, S. Shin, L. Benoit, R. G. Oberprieler, B. D. Farrell, S. D. J. Brown, R. A. B. Leschen, G. J. Kergoat & D. D. McKenna. 2023. Phylogenomics illuminates the phylogeny of flower weevils (Curculioninae) and reveals ten independent origins of brood-site pollination mutualism in true weevils. Proceedings of the Royal Society of London Series B—Biological Sciences 290: 20230889.

[LC40] Laporte de Castelnau, F. 1840. Histoire Naturelle des Insectes Coléoptères vol. 2. P. Duménil: Paris.

[LB20] Letsch, H., M. Balke, E. F. A. Toussaint & A. Riedel. 2020. Historical biogeography of the hyperdiverse hidden snout weevils (Coleoptera, Curculionidae, Cryptorhynchinae). Systematic Entomology 45: 312–326.

[LE23] Li, Y.-D., M. S. Engel, E. Tihelka & C. Cai. 2023. Phylogenomics of weevils revisited: data curation and modelling compositional heterogeneity. Biology Letters 19: 20230307.

[Mac86] Macleay, W. 1886. The insects of the Fly River, New Guinea, “Coleoptera” (continued). Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 183–204.

[ML05] Marvaldi, A. E., & A. A. Lanteri. 2005. Key to higher taxa of South American weevils based on adult characters (Coleoptera, Curculionoidea). Revista Chilena de Historia Natural 78: 65–87.

[Mas86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part V. Proceedings of the Linnean Society of New South Wales, series 2, 1 (3): 585–686.

[M94] May, B. M. 1994. An introduction to the immature stages of Australian Curculionoidea. In: Zimmerman, E. C. Australian Weevils (Coleoptera: Curculionoidea) vol. 2. Brentidae, Eurhynchidae, Apionidae and a chapter on immature stages by Brenda May pp. 365–728. CSIRO Australia.

[PLB13] Poinar, G., Jr, A. A. Legalov & A. E. Brown. 2013. Brachycamacina, a new subtribe of the tribe Naupactini (Coleoptera: Curculionidae Entiminae) in Dominican amber. Palaeontologica Electronica 16 (3): 23A.

[P66] Pascoe, F. P. 1866. Catalogue of longicorn Coleoptera collected in the island of Penang by James Lamb, Esq. Part I. Proceedings of the Zoological Society of London 1866: 222–267.

[W41] Waterhouse, G. R. 1841. Descriptions of some new coleopterous insects from the southern parts of S. America, collected by C. Darwin, Esq., and T. Bridges, Esq. Proceedings of the Zoological Society of London 9: 105–128.

[Z93] Zimmerman, E. C. 1993. Australian Weevils (Coleoptera: Curculionoidea) vol. 3. Nanophyidae, Rhynchophoridae, Erirhinidae, Curculionidae: Amycterinae, literature consulted. CSIRO Australia.

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