Ananias alticonica, from Pan & Erwin (2002).

Belongs within: Vetigastropoda.

The Eotomariinae are a group of gastropods known from the Middle Ordovician to Permian.

Characters (from Knight et al. 1960): Labral slit moderate to deep, umbilicus narrow or absent; ornament collabral and spiral elements, collabral dominant.

    |--Deseretospira Gordon & Yochelson 1987 [Deseretospirides, Deseretospirini]BR05
    |    `--*D. monilifera Gordon & Yochelson 1987BR17
    |--Glabrocingulum Thomas 1940 [Glabrocingulides, Glabrocingulini]BR05
    |    `--*G. beggi Thomas 1940KC60
    `--Eotomariini [Eotomariides, Liospirinae]BR05
         |--Pseudobaylea Dickins 1963W63
         |    `--*P. freneyensis Dickins 1963 [=Mourlonia (Pseudobaylea) freneyensis]W63
         |--Arastra Stoyanow 1948KC60
         |    `--*A. torquata Stoyanow 1948KC60
         |--Bembexia Oehlert & Oehlert 1888FB04
         |    |--*B. larteti (Munier-Chalmas 1876) [=Pleurotomaria larteti]KC60
         |    `--B. sulcomarginataB12
         |--Eotomaria Ulrich & Scofield 1897BR05 [incl. Spiroraphe Perner 1907KC60]
         |    |--*E. sublaevis Ulrich 1897BR17
         |    `--E. canalifera Ulrich in Ulrich & Scofield 1897KC60
         |--Liospira Ulrich & Scofield 1897BR05 [incl. Eocryptaenia Koken 1925KC60]
         |    |--*L. micula (Hall 1862) [=Pleurotomaria micula]KC60
         |    `--L. angustata Ulrich & Scofield 1897YN83
         |--Trepospira Ulrich & Scofield 1897 [incl. Kansana Tasch 1953]KC60
         |    |--*T. sphaerulata (Conrad 1842) [=Pleurotomaria sphaerulata]KC60
         |    |--T. kansuensisG31
         |    `--T. tumidaG31
         |--Clathrospira Ulrich & Scofield 1897KC60
         |    |--*C. subconica (Hall 1847) [=Pleurotomaria subconica]KC60
         |    |--C. angulata Yu & Ning 1983YN83
         |    |--C. conicaYN83
         |    `--C. obscura (Hynda 1986)D94
         `--Ananias Knight 1945W63, PE02
              |--*A. welleri (Newell 1935)PE02 [=Phanerotrema welleriPE02, Glabrocingulum (*Ananias) welleriKC60]
              |--A. alticonica (Wang in Wang & Xi 1980) [=Guizhouspira alticonica]PE02
              |--A. campbelli (Waterhouse 1963) [=Glabrocingulum (Ananias) campbelli]W63
              |--A. gibber Chronic 1952W63
              |--A. guojiashanensis Tong & Erwin 2001TE01
              |--A. johannisaustriae (Klipstein 1843) (see below for synonymy)TE01
              `--A. wannense (Newell 1935)W63

Ananias johannisaustriae (Klipstein 1843) [=Pleurotomaria johannisaustriae, A. joannisaustriae (l. c.); incl. Worthenia apunctata Kittl 1894]TE01

*Type species of generic name indicated


[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[B12] Brett, C. E. 2012. Coordinated stasis reconsidered: a perspective at fifteen years. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 23–36. Springer.

[D94] Dzik, J. 1994. Evolution of ‘small shelly fossils’ assemblages of the early Paleozoic. Acta Palaeontologica Polonica 39 (3): 247–313.

[FB04] Frýda, J., & R. B. Blodgett. 2004. New Emsian (Late Early Devonian) gastropods from Limestone Mountain, Medfra B-4 quadrangle, west-central Alaska (Farewell Terrane), and their paleobiogeographic affinities and evolutionary significance. Journal of Paleontology 78 (1): 111–132.

[G31] Grabau, A. W. 1931. The Permian of Mongolia: A report on the Permian fauna of the Jisu Honguer limestone of Mongolia and its relations to the Permian of other parts of the world. American Museum of Natural History: New York.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[PE02] Pan, H.-Z., & D. H. Erwin. 2002. Gastropods from the Permian of Guangxi and Yunnan Provinces, south China. Journal of Paleontology 76 (Memoir 56): 1–49.

[TE01] Tong, J., & D. H. Erwin. 2001. Triassic gastropods of the southern Qinling Mountains, China. Smithsonian Contributions to Paleobiology 92: 1–47.

[W63] Waterhouse, J. B. 1963. Permian gastropods of New Zealand. Part 3—Pleurotomariacea (concluded). New Zealand Journal of Geology and Geophysics 6 (4): 587–622.

[YN83] Yu W. & Ning H. 1983. Middle Ordovician gastropods from Ruoqiang, Xinjiang. Acta Palaeontologica Sinica 22 (2): 199–203.

Leave a comment

Your email address will not be published. Required fields are marked *