Epichrysomallidae

 Meselatus sp., from Biodiversity Institute of Ontario.

Belongs within: Chalcidoidea.

The Epichrysomallidae are a Palaeotropical group of wasps that oviposit into figs, with the larvae feeding on ovary tissues within the fig and transforming them into a gall (Bouček 1988).

Characters (from Bouček 1988, as Epichrysomallinae): Body non-metallic. frequently yellowish, reddish or brown to blackish; usually without dense and deep sculpture (except for Asycobia), without regular dense pilosity but generally with only spaced paired setae on head and thorax, very rarely with some more hairs in deeper parts and on sides. Occiput primarily with occipital carina but in several genera this is obliterated or absent. C1ypeus delimited at least on sides, often also supraclypeal area well defined; lower clypeal margin produced, usually bilobed. Antennae mostly near to centre of face, usually close together. Antennae rarely 13-segmented, often reduced in number of segments when first flagellar segment, reduced to anellus, fuses with the second, or the second is reduced in size and then fuses with the next segment. In extreme case (males of Camarothorax) the whole flagellum is reduced to 2 or 3 segments (and wings also reduced). Antennal formula often different in the two sexes, half-fused segments in one antenna commonly occur. Thorax in lateral view often convex, sometimes elongate, especially in males of some species. Pronotal collar almost never defined. Scutum with notauli primarily complete but in some species not so. Scutellum without frenal groove, without distinct axillar grooves, convex, with admarginal groove along apex. Propodeum fairly large, usually without median carina and without plicae but sometimes with coarser sculpture medially, widening along anterior margin and in sulcus behind spiracle. Metapleuron broad, tapering to base of hindwing. Mesepimeron separated by irregular groove; mesopleuron with sculptured femoral depression. Prepectus large, triangular. Legs of medium length; apex of fore and hind tibiae oblique at spurs, hence these moved slighty away from apex; two hind spurs; hind basitarsus at least as long as following two segments combined, in some species fourth segment fused with the fifth, hence tarsi 4-segmented. Hind coxa attached very high. Wing pilosity more or less reduced; postmarginal vem almost always shorter than the stigmal, sometimes quite rudimentary; stigmal vein fairly long, arising at very nearly a right angle, often curved; not distinctly knobbed. Gaster high, often compressed from side to side, hind margins of basal 2 or 3 tergites usually excised as an adaptation to help coiling in (and out) of the relatively long ovipositor. However, sheaths not distinctly exserted. Hypopygium large, ending usually in posterior third of gaster. Petiole in some males conspicuous, rarely also in females. Cerci usually slightly elongate, placed inside of epipygium, this as one sclerite (fused tergites 7 and 8).

<==Epichrysomallidae [Epichrysomallinae]B88
    |--Neosycophila Grandi 1923MH11, B88
    |    `--*N. omeomorphaB88
    |--Acophila Ishii 1934MH11, B88
    |    `--*A. mikiiB88
    `--+--+--Eufroggattisca Ghesquière 1946MH11, B88 (see below for synonymy)
       |  |    `--*E. polita (Ashmead 1904) [=*Froggattia polita, *Eufroggattia polita, *Eufroggattiana polita]B88
       |  `--+--Epichrysomalla Girault 1915MH11, B88
       |     |    `--*E. atricorpus Girault 1915B88
       |     `--Camarothorax Mayr 1906MH11, B88 [incl. Pilkhanivora Farooqi & Ramdas Menon 1973B88, Sycobiomorpha Joseph 1961B88]
       |          |--*C. obscurusB88
       |          `--C. bimasculinus [=*Sycobiomorpha bimasculina; incl. *Pilkhanivora trimorpha]B88
       `--+--SycophilomorphaMH11
          `--+--SycobiaMH11
             `--+--Herodotia Girault 1931MH11, B88
                |    |--*H. procopii Girault 1931B88
                |    |--‘Eufroggattia’ okinawensis Ishii 1934B88
                |    `--H. subatriventris (Girault 1923) [=Meselatus subatriventris]B88
                `--+--SycotetraMH11
                   `--+--LachaiseaMH11
                      `--Meselatus Girault 1922MH11, B88 (see below for synonymy)
                           |--*M. ficus Girault 1922B88
                           |--M. fasciatipennis Girault 1929B88
                           `--M. leai (Dodd 1924) (see below for synonymy)B88
Epichrysomallidae incertae sedis:
  Asycobia Bouček 1988B88
    `--*A. punctulata Bouček 1988B88
  Odontofroggatia Ishii 1934B88
    |--*O. gajimaruB88
    |--O. corneri Wiebes 1980B88
    |--O. galili Wiebes 1980B88
    `--O. ishii Wiebes 1980B88
  JosephiellaBM22
  LeeuweniellaBM22
  ParasycobiaBM22
  SycobiomorphellaBM22
  SycomacophilaBM22
  SycophilodesBM22

Eufroggattisca Ghesquière 1946MH11, B88 [=Eufroggattia Ashmead 1904 non Goding 1903B88, Eufroggattiana Gahan in Gahan & Ferrière 1947B88, Froggattia Ashmead 1904 non Horvarth 1902B88]

Meselatus Girault 1922MH11, B88 [incl. Austroperilampus Dodd 1924B88, Pembertonia Gahan in Gahan & Ferriére 1947 non Zimmerman 1944B88]

Meselatus leai (Dodd 1924) [=*Austroperilampus leai; incl. *Pembertonia ficicola Gahan in Gahan & Ferriére 1947]B88

*Type species of generic name indicated

References

[B88] Bouček, Z. 1988. Australasian Chalcidoidea (Hymenoptera): A biosystematic revision of genera of fourteen families, with a reclassification of species. CAB International: Wallingford (UK).

[BM22] Burks, R., M.-D. Mitroiu, L. Fusu, J. M. Heraty, P. Janšta, S. Heydon, N. D.-S. Papilloud, R. S. Peters, E. V. Tselikh, J. B. Woolley, S. van Noort, H. Baur, A. Cruaud, C. Darling, M. Haas, P. Hanson, L. Krogmann & J.-Y. Rasplus. 2022. From hell’s heart I stab at thee! A determined approach towards a monophyletic Pteromalidae and reclassification of Chalcidoidea (Hymenoptera). Journal of Hymenoptera Research 94: 13–88.

[MH11] Munro, J. B., J. M. Heraty, R. A. Burks, D. Hawks, J. Mottern, A. Cruaud, J.-Y. Rasplus & P. Jansta. 2011. A molecular phylogeny of the Chalcidoidea (Hymenoptera). PloS One 6 (11): e27023.

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