Euarthropoda

 Reconstruction of Mollisonia plenovenatrix by J. Liang, from Aria & Caron (2019).

Belongs within: Arthropoda.
Contains: Pycnogonida, Euchelicerata, Artiopoda, Marrellomorpha, Myriapoda, Hymenocarina, Euthycarcinoidea, Oligostraca, Allotriocarida, Tantulocarida, Thecostraca, Copepoda, Malacostraca.

The name Euarthropoda refers to the most exclusive clade formed by crown-group arthropods. Recent analyses agree on a division of living arthropods between three major clades, the Chelicerata, Myriapoda and Pancrustacea, with the latter two usually being united to form a clade of arthropods possessing mandibles. Fossil euarthropod taxa not belonging to these clades are best known from the Cambrian period with the exception of the Artiopoda (the clade including the trilobites). The Chelicerata have a single pair of pre-oral appendages, the chelicerae, that are usually adapted for manipulating food.

The Cambrian Mollisoniidae are a group of likely stem-chelicerates characterised by a broad, helmet-like head shield bearing wide eye notches, seven free segments in the anterior portion of the trunk extending into short pleurae, and posterior trunk segments fused into a pygidium (Aria & Caron 2019). The Middle Cambrian genus Sanctacaris has a biramous pre-oral limb, raptorial post-oral head limbs, and a flabellate telson (Bousfield 1995). Another Cambrian group, the Retifaciida, are characterised by a short, broad head shield, a large, oval pygidium and an elongate, multisegmented telson).

The clade Pancrustacea includes the modern crustaceans and insects with the latter a derived subgroup of the former (Regier et al. 2010). This last grouping is primarily supported by molecular data; potential synapomorphies of the Pancrustacea have all become modified in various subclades. Relationships within the pancrustaceans remain subject to analysis. A relationship between the Tantulocarida and Thecostraca has been suggested on the basis of gonopore position in the two groups; tantulocarids and basal thecostracans are also both sessile parasites.

<==Euarthropoda (see below for synonymy)
    |  i. s.: Dala peilertae Müller 1983ZS07
    |         Walossekia quinquespinosa Müller 1983ZS07
    |         Hexona Risso 1826R26
    |           `--*H. parasitica Risso 1826R26
    |         Zuphea Risso 1826R26
    |           `--*Z. sparicola Risso 1826R26
    |--PanchelicerataAC19
    |    |--+--Sanctacaris Briggs & Collins 1988AC19, CB04 [Sanctacarida, Sanctacaridea]
    |    |  |    |--‘Utahcaris’ orion Conway Morris & Robison 1988B95
    |    |  |    `--Sanctacaris uncata Briggs & Collins 1988WD02
    |    |  `--HabeliidaB95
    |    |       |--EcnomocarisLSE13
    |    |       `--Habelia Walcott 1912AC19, CB04
    |    |            `--H. optataAC19
    |    `--ChelicerataAC19
    |         |  i. s.: Wisangocaris barbarahardyae Jago, García-Bellido & Gehling 2016BS20
    |         |--PycnogonidaRS10
    |         |--EuchelicerataRS10
    |         `--Mollisoniidae [Mollisoniida]AC19
    |              |--Thelxiope Simonetta & Delle Cave 1975AC19
    |              |--Urokodia Hou, Chen & Lu 1989AC19
    |              |--Corcorania Jell 1980AC19
    |              |    `--C. trispinosaAC19
    |              `--Mollisonia Walcott 1912AC19, CB04 [incl. Houghtonites Raymond 1931AC19]
    |                   |--*M. symmetrica Walcott 1912 [incl. M. rara Walcott 1912]AC19
    |                   |--M. gracilis Walcott 1912 [=Houghtonites gracilis]AC19
    |                   |--M. plenovenatrix Aria & Caron 2019AC19
    |                   `--M. sinica Zhang et al. 2002AC19
    `--+--RetifaciidaLSE13
       |    |--Retifacies Hou, Chen & Lu 1989LSE13, CB04
       |    |    `--R. abnormalis Hou et al. 1989CB04
       |    `--+--PygmaclypeatusLSE13
       |       `--+--+--KiisortoqiaAC19
       |          |  `--SiriocarsLSE13
       |          `--SquamaculaLSE13
       |               |--S. buckorumLSE13
       |               `--S. clypeatusLSE13
       `--+--ArtiopodaAC19
          `--+--MarrellomorphaAC19
             `--+--Aquilonifer Briggs, Siveter et al. 2016AC19, BS16
                |    `--*A. spinosus Briggs, Siveter et al. 2016BS16
                `--Mandibulata [Antennata, Atelocerata, Biantennata, Labiophora, Labrophora, Opisthogoneata, Tracheata]AC17
                     |  i. s.: Ercaia [Promandibulata]C12
                     |           `--E. minusculeC12
                     |         MusacarisLSE13
                     |         CambropachycopidaeLSE13
                     |           |--Cambropachycope clarksoni Walossek & Müller 1990BWW93
                     |           `--Goticaris longispinosa Walossek & Müller 1990BWW93
                     |         Devonopilio Tihelka, Tian & Cai 2020P-GS21, TTC20
                     |           `--*D. hutchinsoni Tihelka, Tian & Cai 2020TTC20
                     |--MyriapodaAC19
                     `--+--+--HymenocarinaAC19
                        |  `--+--EuthycarcinoideaAC19
                        |     `--Fuxianhuiida [Fuxianhuiata, Proarthropoda]LSE13
                        |          |  i. s.: LiangwangshaniaLSE13
                        |          |--+--Shankouia zhengheiAC19, C12
                        |          |  `--ChengjiangocarisAC19
                        |          |       |--C. kunmingensis Yang, Ortega-Hernández et al. 2013YO-H13
                        |          |       `--C. longiformis Hou & Bergström 1991B95
                        |          `--+--GuangweicarisLSE13
                        |             `--FuxianhuiaAC19
                        |                  |--F. protensa Hou 1987B95
                        |                  `--F. xiaoshibaensis Yang, Ortega-Hernández et al. 2013YO-H13
                        `--Pancrustacea (see below for synonymy)AC17
                             |  i. s.: Nippotantulus heteroxenus Huys, Ohtsuka & Boxshall 1994MA01
                             |         NeomysisM02
                             |           |--N. americanaM02
                             |           `--N. vulgarisCS77
                             |         Perosquilla armataK-MC02
                             |           |--P. a. armataK-MC02
                             |           `--P. a. capensisK-MC02
                             |         Mesosoma Otto 1821C92
                             |         Mixodiaptomus laciniatusD51
                             |         Pleuroncodes planipesCH97
                             |         Phalangites priscus Münster 1836D07
                             |         Cryptophthalmus Rafinesque 1814BR05
                             |         Cymodocea Rafinesque 1814BR05
                             |         ‘Cymodocea’ Leach 1818 nec Rafinesque 1814 nec Lamouroux 1816BR05
                             |         ‘Dactylopus’ Claus 1862 non Gill 1859BR05
                             |         Donovania Leach 1814BR05
                             |         Orientalina Kolosnitsyna 1973BR05
                             |         Pagodina van Beneden 1853BR05
                             |         ‘Trichia’ de Haan 1839 non Haller 1768BR05
                             |         Fenneropenaeus chinensisDH08
                             |         Limnadella kitei Girard 1854DS89
                             |         Anatanais robustoBBB-S95
                             |         Anthonema Walther 1904H75
                             |           `--*A. problematica Walther 1904H75
                             |         ‘Acherusia’ Lucas 1846 non Laporte & Gory 1837E12
                             |           `--A. dumerilii Lucas 1846E12
                             |         Naesea edwardsii Lucas 1846E12
                             |         Ovalia Latreille in Griffith & Pidgeon 1833KA-Z11
                             |         CycloideaFGS04
                             |           |--Cyclus de Koninck 1841 [incl. Paraprosopon Gemmellaro 1890]FGS04
                             |           `--Oonocarcinus Gemmellaro 1890FGS04
                             |         HeymonsicambriaEL11
                             |         Hymenodora glacialisCS77
                             |         Ozolus Latreille 1802L02a
                             |           `--*O. gasterostei Latreille 1802L02b (see below for synonymy)
                             |         AmymoneL02b
                             |           |--A. baccha [=Monoculus bacchus]L02b
                             |           |--A. faunus [=A. fauna, Monoculus faunus]L02b
                             |           |--A. moenasL02b
                             |           |--A. satyrus [=A. satyra, Monoculus satyrus]L02b
                             |           |--A. silene [=A. silenus, Monoculus silenus]L02b
                             |           `--A. thyasL02b
                             |         NaupliusL02b
                             |           |--N. bracteatus [=Monoculus bracteatus]L02b
                             |           `--N. saltatorius [=Monoculus saltatorius]L02b
                             |         Microdentopus gryllotalpaPP64
                             |         Aristeomorpha foliaceaPP64
                             |         Ligur ensiferusPP64
                             |         Paromola cuvieriPP64
                             |         Anamathia rissoanaPP64
                             |         Ergasticus cloueiPP64
                             |         Dorynchus thompsoniPP64
                             |         Cymopolia caroniPP64
                             |         Halopsyche de Saussure 1857BR17
                             |         Palpipes Roth 1851K18
                             |         Berndtia purpureaCG97
                             |         Trypetasa lampasCG97
                             |         Amblyops crozetiiO01
                             |         Harpactocarcinus tumidusH79
                             |         TuzoiidaeO68
                             |           |--TuzoiaO68
                             |           `--*Dioxycaris argenta (Walcott 1888) [=Leperditia argenta, Isoxys argenta]O68
                             |         Corynuropsis Scott 1894E00
                             |         Cytherea spinosaR26
                             |--OligostracaAC19
                             `--Altocrustacea [Ostraca, Phyllopodomorpha, Thoracopoda]RS10
                                  |--AllotriocaridaSC17
                                  `--Multicrustacea [Communostraca, Vericrustacea]SC17
                                       |  i. s.: Arenosicaris inflataSC17
                                       |--+--TantulocaridaLS02
                                       |  `--ThecostracaSC17
                                       `--+--CopepodaSC17
                                          `--MalacostracaSC17

Euarthropoda [Arachnata, Arachnomorpha, Aspidonia, Aspidota, Cormogonida, Eotrilobitacea, Myriochelata, Paradoxopoda, Polymeria, Skaracarida]

*Ozolus gasterostei Latreille 1802L02b [incl. Argulus delphiniL02b, Monoculus delphinusL02a, M. gyriniL02b]

Pancrustacea [Archoptera, Copepodoida, Copepodomorpha, Crustacea, Crustaceanomorpha, Entomostraca, Eucrustacea, Gymnota, Maxillopoda, Pneumonura, Poecilopoda, Progonomorpha, Pseudopoda, Tetraconata, Thecostracomorpha]AC17

*Type species of generic name indicated

References

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