Cyphoderia ampulla, copyright Picturepest.

Belongs within: Imbricatea.
Contains: Psammonobiotidae, Euglyphidae.

The Euglyphida are a group of testate amoeboids, often with a covering of silica scales (Adl et al. 2012).

Characters (from Adl et al. 2012): Test of organic material; usually with secreted silica scales held together by organic cement; tubular mitochondrial cristae.

Euglyphida [Euglyphacea]
    |  i. s.: MatsakisionAS12
    |         PsammonobiotidaeAS12
    |    |--Tracheleuglypha Deflandre 1928C-SC03, LT64
    |    |    |--*T. dentata (Vejdovský 1882) [=Euglypha dentata]LT64
    |    |    `--T. acollaLNB03
    |    `--+--AssulinidaeAS12
    |       |    |--ValkanoviaAS12
    |       |    |--Placocista Leidy 1879AS12, LT64 [=Placocysta Blochmann 1886LT64]
    |       |    |    `--*P. spinosa (Carter 1865) [=Euglypha spinosa, *Placocysta spinosa]LT64
    |       |    `--Assulina Ehrenberg 1872C-SC03, LT64
    |       |         |--*A. seminulum (Ehrenberg 1848) [=Difflugia seminulum, Sphenoderia seminulum]LT64
    |       |         `--A. muscorumC-SC03
    |       `--+--+--EuglyphidaeBM05
    |          |  `--SphenoderiidaeAB19
    |          |       |--Trachelocorythion pulchellumC-SC03
    |          |       |--DeharvengiaAB19
    |          |       `--Sphenoderia Schlumberger 1845 [=Hologlypha Ehrenberg 1872]LT64
    |          |            `--*S. lenta Schlumberger 1845 [=Difflugia (Assulina) lenta, *Hologlypha lenta]LT64
    |          `--Trinematidae [Trinematinae, Trinemidae, Trineminae]LT64
    |               |--PlayfairinaAS12
    |               |--PuytoraciaAS12
    |               |--Corythion Taránek 1882LT64
    |               |    |--*C. dubium Taránek 1882LT64
    |               |    |--C. delmareiLB90
    |               |    `--C. pulchellumV63
    |               `--Trinema Dujardin 1841BM05, LT64 [incl. Homoeochlamys Ehrenberg 1872LT64]
    |                    |--T. enchelys (Ehrenberg 1838) (see below for synonymy)LT64
    |                    |--T. complanatum [incl. T. complanatum var. inaequalis]LNB03
    |                    `--T. lineareG86
    `--+--Paulinellidae [Paulinellinae]AS12
       |    |--OvulinataAS12
       |    |--MicropyxidiellaAB19
       |    `--Paulinella Lauterborn 1895BM05, MM05 [incl. Cyanospira Chodat 1920LT64]
       |         |--*P. chromatophora Lauterborn 1895MM05
       |         |--*Cyanospira’ aeruginosa Chodat 1920LT64
       |         `--P. ovalisMM05
       `--Cyphoderiidae [Revolventiidae]N03
            |--Messemvriella Golemansky 1973N03, G86
            |    `--M. filosa Golemansky 1973G86
            |--Pseudocyphoderia Sudzuki 1979G86
            |    `--P. orientalis Sudzuki 1979G86
            |--Schaudinnula Averintsev 1906AS12, LT64
            |    `--*S. arcelloides Averintsev 1906 [=Cyphoderia arcelloides]LT64
            |--Corythionella Golemansky 1970N03, G86
            |    |--C. acolla Golemansky 1970G86
            |    |--C. anteroplanata Chardez 1977G86
            |    |--C. minima Golemansky 1970 [incl. C. minima var. nipponica Sudzuki 1979]G86
            |    |--C. pontica Golemansky 1970G86
            |    `--C. sudzukii Chardez 1977G86
            |--Pseudocorythion Valkanov 1970N03, G86
            |    |--P. acutum (Wailes 1927) [incl. P. acutum var. nipponicum Sudzuki 1979]G86
            |    |--P. mannei Chardez 1971G86
            |    |--P. nipponicum Sudzuki 1976G86
            |    |--P. undulacollis Chardez & Thomas 1980G86
            |    `--P. wailesi Golemansky 1971G86
            `--Cyphoderia Schlumberger 1845G86 (see below for synonymy)
                 |--C. ampulla (Ehrenberg 1840) (see below for synonymy)LT64
                 |    |--C. a. ampullaN03
                 |    `--C. a. bicornisN03
                 |--C. compressa Golemansky 1979G86
                 |--C. laevisN03
                 |--‘Difflugia’ lagena Ehrenberg 1843 [=*Allodictya lagena, D. (Reticella) lagena]LT64
                 |--C. littoralis Golemansky 1973 [incl. C. littoralis var. simodensis Sudzuki 1979]G86
                 `--C. lunata Štĕpánek 1967N03

Cyphoderia Schlumberger 1845G86 [incl. Allodictya Ehrenberg 1872LT64, Ampullaria Werneck in Ehrenberg 1872 non Lamarck 1799LT64]

Cyphoderia ampulla (Ehrenberg 1840) [=Difflugia ampulla, *Ampullaria ampulla; incl. *Cyphoderia margaritacea Schlumberger 1845]LT64

Trinema enchelys (Ehrenberg 1838) [=Difflugia enchelys, Arcella (*Homoeochlamys) enchelys; incl. *Trinema acinus Dujardin 1841]LT64

*Type species of generic name indicated


[AB19] Adl, S. M., D. Bass, C. E. Lane, J. Lukeš, C. L. Schoch, A. Smirnov, S. Agatha, C. Berney, M. W. Brown, F. Burki, P. Cárdenas, I. Čepička, L. Chistyakova, J. del Campo, M. Dunthorn, B. Edvardsen, Y. Eglit, L. Guillou, V. Hampl, A. A. Heiss, M. Hoppenrath, T. Y. James, A. Karnkowska, S. Karpov, E. Kim, M. Kolisko, A. Kudryavtsev, D. J. G. Lahr, E. Lara, L. Le Gall, D. H. Lynn, D. G. Mann, R. Massana, E. A. D. Mitchell, C. Morrow, J. S. Park, J. W. Pawlowski, M. J. Powell, D. J. Richter, S. Rueckert, L. Shadwick, S. Shimano, F. W. Spiegel, G. Torruella, N. Youssef, V. Zlatogursky & Q. Zhang. 2019. Revisions to the classification, nomenclature, and diversity of eukaryotes. Journal of Eukaryotic Microbiology 66: 4–119.

[AS12] Adl, S. M., A. G. B. Simpson, C. E. Lane, J. Lukeš, D. Bass, S. S. Bowser, M. W. Brown, F. Burki, M. Dunthorn, V. Hampl, A. Heiss, M. Hoppenrath, E. Lara, E. Le Gall, D. H. Lynn, H. McManus, E. A. D. Mitchell, S. E. Mozley-Stanridge, L. W. Parfrey, J. Pawlowski, S. Rueckert, L. Shadwick, C. L. Schoch, A. Smirnov & F. W. Spiegel. 2012. The revised classification of eukaryotes. Journal of Eukaryotic Microbiology 59 (5): 429–493.

[BM05] Bass, D., D. Moreira, P. López-García, S. Polet, E. E. Chao, S. von der Heyden, J. Pawlowski & T. Cavalier-Smith. 2005. Polyubiquitin insertions and the phylogeny of Cercozoa and Rhizaria. Protist 156: 149–161.

[C-SC03] Cavalier-Smith, T., & E. E.-Y. Chao. 2003. Phylogeny and classification of phylum Cercozoa. Protist 154: 341–358.

[G86] Golemansky, V. G. 1986. Rhizopoda: Testacea. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 5–16. E. J. Brill/Dr. W. Backhuys: Leiden.

[LNB03] Ledeganck, P., I. Nijs & L. Beyens. 2003. Plant functional group diversity promotes soil protist diversity. Protist 154 (2): 239–249.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[LB90] Lousier, J. D., & S. S. Bamforth. 1990. Soil protozoa. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 97–136. John Wiley & Sones: New York.

[MM05] Melkonian, M., & D. Mollenhauer. 2005. Robert Lauterborn (1869–1952) and his Paulinella chromatophora. Protist 156: 253–262.

[N03] Nicholls, K. H. 2003. Form variation in Campascus minutus and a description of Campascus simcoei sp. n. (Testaceafilosea, Psammonobiotidae). European Journal of Protistology 39 (1): 103–112.

[V63] Varga, L. 1963. Weitere Untersuchungen über die aquatile Mikrofauna der Baradla-Höhle bei Aggtelek (Ungarn) (Biospeologica Hungarica, XVII). Acta Zoologica Academiae Scientiarum Hungaricae 9 (3–4): 439–458.

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