Counterpart, part and interpretive diagram of holotype of Saxonagrion minutus, from Nel et al. (1999).

Belongs within: Pandiscoidalia.
Contains: Zygoptera, Euepiproctophora, Isophlebioptera.

The Euodonata is the crown group of dragonflies and damselflies. Synapomorphies of this clade include the formation of the posterior arculus at the separation of veins RP and MA, and the presence of the subdiscoidal vein (Grimaldi & Engel 2005). The earliest record of this clade is from the Late Permian, with the stem-damselfly Saxonagrion minutus being found in the Salagou Formation of France (Nel et al. 1999).

    |  `--SaxonagrionNB01 [SaxonagrionidaeGE05]
    |       `--S. minutus Nel et al. 1999NB01
    `--Epiprocta [Epiproctophora]GE05
         |  i. s.: OreopteridaeGE05
         |         TuranothemistidaeGE05
            `--SteleopteronRJ93 [SteleopteridaeRP02]
                 `--S. deichmuelleriRJ93
Euodonata incertae sedis:
  Shurabiola Pritykina 1980FN02
  Mortonagrion hirosei Asahina 1972I92
    |--R. aculeataI92
    |    |--R. a. aculeataI92
    |    |--R. a. hiraoiI92
    |    `--R. a. yakusimensisI92
    `--R. okinawanus Asahina 1951I92
  Boninthemis insularis (Matsumura 1913)I92
    |--L. elegantissimaI92
    `--L. tricolor Ris 1916I92
    |--A. amamiensis (Asahina 1962)I92
    |    |--A. a. amamiensisI92
    |    `--A. a. okinawanus (Asahina 1964)I92
    `--A. yayeyamensis (Oguma 1926)I92
    |--L. sauteriI92
    |    |--L. s. sauteriI92
    |    `--L. s. formosanusI92
    `--L. yayeyamensis Oguma 1926I92
  Macromidia ishidai Asahina 1964I92
  Cercion calamorumI92
  Boninagrion ezoin Asahina 1952I92
    |--S. ryukyuanusI92
    |    |--S. r. ryukyuanusI92
    |    `--S. r. asatoi Asahina 1972I92
    `--S. shirozuiI92
         |--S. s. shirozuiI92
         `--S. s. watanabei Asahina 1984I92
    |--P. ishigakiana Asahina 1951I92
    |    |--P. i. ishigakianaI92
    |    `--P. i. nagaminei Asahina 1988I92
    `--P. risiI92
         |--P. r. risiI92
         `--P. r. sakishimana Asahina 1964I92
  Erythrodiplax berenice (Drury 1770)SB05

*Type species of generic name indicated


[FB03] Fleck, G., G. Bechly, X. Martínez-Delclòs, E. Jarzembowski, R. Coram & A. Nel. 2003. Phylogeny and classification of the Stenophlebioptera (Odonata: Epiproctophora). Annales de la Societe Entomologique de France 39 (1): 55–93.

[FN02] Fleck, G., & A. Nel. 2002. The first isophlebioid dragonfly (Odonata: Isophlebioptera: Campterophlebiidae) from the Mesozoic of China. Palaeontology 45 (6): 1123–1136.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[NB01] Nel, A., O. Bethoux, G. Bechly, X. Martínez-Delclòs & F. Papier. 2001. The Permo-Triassic Odonatoptera of the “protodonate” grade (Insecta: Odonatoptera). Annales de la Société Entomologique de France (N.S.) 37 (4): 501–525.

[RP02] Rasnitsyn, A. P., & L. N. Pritykina. 2002. Superorder Libellulidea Laicharting, 1781. Order Odonata Fabricius, 1792. The dragonflies. In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 97–104. Kluwer Academic Publishers: Dordrecht.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[SB05] Sala, J., & D. Boix. 2005. Presence of the Nearctic water boatman Trichocorixa verticalis verticalis (Fieber, 1851) (Heteroptera, Corixidae) in the Algarve region (S Portugal). Graellsia 61 (1): 31–36.

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