Pisinna zosterophila, copyright Eddie Hardy.

Belongs within: Mollusca.
Contains: Scoliostomatidae, Caseolus, Leiostyla, Clisospiroidea, Bellerophontoidea, Archinacelloidea, Pelagielloidea, Macluritida, Neomphaliones, Patellogastropoda, Vetigastropoda, Neritimorpha, Loxonematoidea, Heterobranchia, Peruneloidea, Cyclophoroidea, Ampullariidae, Viviparoidea, Orthonematoidea, Subulitoidea, Soleniscidae, Meekospiridae, Cerithiimorpha, Hypsogastropoda.

The Gastropoda, snails and slugs, are a major clade of molluscs characterised by the process of torsion during larval development with the visceral mass and mantle becoming turned 180° relative to the head and foot. The shell (if present) has a single valve and is often spirally coiled; an operculum is also often present that closes the aperture of the shell when the body is withdrawn. Both of these features may be modified in derived subgroups: the shell may be lost or coiling reduced, and torsion may be secondarily reversed over the course of continued development. Gastropods are ancestrally aquatic but a number of derived lineages have become terrestrial. Embryonic development in aquatic gastropods may be indirect, with a free-living, planktotrophic larval stage, or direct, with lecithotrophic embryos hatching into crawling juveniles. These differing modes of development are often reflected in different aspects of the protoconch (the initial, larval section of the shell which often differs in appearance from the mature teleoconch) with planktotrophic developers have more whorls to the protoconch than lecithotrophs.

Historically, gastropods have often been divided between three major groupings—the Prosobranchia (possessing a gill and torted anatomy at maturity), Opisthobranchia (gilled but with torsion not evident at maturity), and Pulmonata (respiring via a lung)—but the ‘prosobranchs’ are recognised as paraphyletic to the last two groups (which together form part of the broader clade Heterobranchia). The ‘prosobranchs’ have in turn been historically divided between the grades Archaeogastropoda, Mesogastropoda and Neogastropoda but, again, the ‘archaeogastropods’ are now regarded as paraphyletic with the Neogastropoda and most ‘mesogastropods’ forming the clade Caenogastropoda. The Caenogastropoda and Heterobranchia together form a clade Apogastropoda, supported by a number of synapomorphies such as the presence of a single columellar muscle, loss of the right auricle, and the oesophagus originating at the posterior end of the buccal mass (Simone 2011).

Various Early Palaeozoic fossils such as Bellerophontoidea and Pelagielloidea resemble modern gastropods in the possession of a coiled shell but have not been universally accepted as gastropods due to uncertainty whether or not they exhibited torsion. Thomas et al. (2020) argued for the presence of torsion in Pelagielloidea on the basis of functional interpretations of preserved chaetae and internal grooves on the shell.

<==Gastropoda (see below for synonymy)
| `--MacluritidaTRM20
| `--VetigastropodaS11
`--Adenogonogastropoda [Branchiata, Cyclostomacea, Ectophthalma, Teleohydrophila]S11
`--Apogastropoda (see below for synonymy)S11
| i. s.: Protostylus Mansuy 1914 (n. d.)KC60
| `--*P. lantenoisi Mansuy 1914KC60
| ‘Cyclora’ imbricata Whiteaves 1881G88
| ‘Cyclora’ valvatiformis Whiteaves 1881G88
| OnisciaR42
| |--O. dennisoni Reeve 1842R42
| |--O. strombiformis Reeve 1842R42
| `--O. tuberculosaC64
`--Caenogastropoda (see below for synonymy)S11
| i. s.: Gaeatrivia millardi Cate 1979M03
| Serratovolva dondani Cate 1964M03
| Decorochilina Frýda 1999F01
| Plicatus Pan & Erwin 2002 [Plicatusidae]BR05
| |--*P. scalaris Pan & Erwin 2002PE02
| `--P. multifilaris Pan & Erwin 2002PE02
| Spanionema Whidborne 1891 [Spanionematidae, Spanionematoidea]BR05
| `--*S. scalaroides (Whidborne 1889) [=Loxonema scalaroides]KC60
| Spirochrysalis Kittl 1894NE02
| `--S. protomicra Pan 1982NE02
| Lampanella minimaPC08
| Pragoscutula Frýda 1998PC08, BR05 [Pragoscutulidae]
| `--P. wareni Frýda 1998FB01
| Dendropupoidea [Anthracopupoidea, Dendropupinae, Procyclophoroida]BR05
| |--Maturipupa Pilsbry 1926KC60
| | `--*M. vermilionensis (Bradley 1872) [=Pupa vermilionensis]KC60
| |--Anthracopupa Whitfield 1881 [Anthracopupidae, Anthracopupinae]BR05
| | |--*A. ohioensis Whitfield 1881KC60
| | `--A. britannica Cox 1926TTE93
| `--Dendropupa Owen 1859BR17 (see below for synonymy)
| |--*D. vetusta (Dawson 1859) [=Pupa vetusta]KC60
| |--D. grandaeva (Dawson 1880)TTE93
| `--D. primaeva (Matthew 1895)TTE93
| Acanthonema Grabau in Sherzer & Grabau 1909BR05 [AcanthonematidaeBR17, Acanthonematinae]
| `--A. newberryi (Meek 1871)TTE93 (see below for synonymy)
| AmpezzanildidaeBR17
| |--Cassianilda Bandel 1995NE02
| |--Stuorilda Bandel 1995NE02
| `--Ampezzanilda Bandel 1994BR05
| `--*A. aialensis (Zardini 1980) [=Promathildia aialensis]BR17
| Kittlidiscus Haas 1953BR05 (see below for synonymy)
| `--K. bronni (Klipstein 1845)KC60 (see below for synonymy)
| Spirostylidae [Spirostylinidae]TE01
| |--Spirostylus Kittl 1894TE01
| | |--*S. subcolumnaris (Münster 1841) [=Melania subcolumnaris]TE01
| | `--S. linctus (Böhm 1895) (see below for synonymy)TE01
| `--ClimacinaTTE93
| |--C. catharinae Gemmellaro 1876TTE93
| `--C. mariae Gemmellaro 1876TTE93
| PeruneloideaBR17
`--Epiarthroidea [Teleobranchia, Vivipariformes, Vivipariformii]S11
`--Sorbeoconcha (see below for synonymy)S11
| i. s.: Globocornus Espinosa & Ortea 2010 [Globocornidae]BR17
| `--*G. darwini Espinosa & Ortea 2010BR17
| Acteonina d’Orbigny 1850BR05 (see below for synonymy)
| |--*A. carbonaria (de Koninck 1843) [=Chemnitzia carbonaria]BR05
| |--A. acuta (d’Orbigny 1841) [=Tornatina acuta]BR17
| `--A. obesa Stoliczka 1868TTE93
| OrthonematoideaBR17
| Subulitina [Subulitata]GS75
| |--SubulitoideaGS75
| `--SoleniscoideaBR17
| |--SoleniscidaeBR17
| |--MeekospiridaeBR17
| `--AnozygidaeBR17
| |--Anozyga Hoare 1980 [Anozyginae]BR05
| | `--*A. bulla Hoare 1980BR17
| `--Tmetonema Longstaff 1912 [Tmetoneminae]BR05
| `--*T. subsulcatum Longstaff 1912KC60
| PluviostillaBR17

Gastropoda incertae sedis:
Serpentubina Horný 1964FBL02
Kiviasukkaan Peel 1975FBL02
Pragoserpulina Frýda 1998 [Pragoserpulinidae]BR05
`--*P. tomasi Frýda 1998BR17
Luoguispira Pan & Erwin 2002PE02
|--*L. micra Pan & Erwin 2002PE02
|--L. lineata Pan & Erwin 2002PE02
`--L. multilinea Pan & Erwin 2002PE02
Luoguella Pan & Erwin 2002PE02
|--*L. elegantula Pan & Erwin 2002 [=Gyrospira elegantula]PE02
`--L. diana Pan & Erwin 2002 [=Gyrospira diana]PE02
Kalchreuthia Gründel & Nützel 1998GK02
Paladilha (Lartella Cossmann 1921)GK02
Pseudotaphrus (Pezantia Cossmann 1896)GK02
Derjuginella rufofasciataGAS03
Erheria vinctaGAS03
Yokoyamia ornatissimaGAS03
Epheria turritaKBC03
Tersiella plicosaKBC03
Ausola angustataKBC03
Hinia cuvieriM62
Neritula donovaniM62
Oxytrema Rafinesque 1819B63
`--O. siliculaB96
Hystrivasum horridum Heilprin 1886HP04
Benthonella tenella (Jeffreys 1869)OT04
Chlupacispira spinosa Blodgett & Rohr 1989FB04
Eulepetopsis vitreaGO06
Purkynespira Frýda & Bandel 1997FB01
`--P. reticulataFB01
Streptopelma henchmani Marwick 1926F26
Buccinulus huttoni Kirk 1882F26
Triploca Tate 1894F26
|--T. ligataF26
`--T. waihaoensis Marshall & Murdoch 1923F26
Sublacuna Cossmann 1899 [=Strebloramphus Tate 1898 (preoc.)]F27
`--*S. mirula (Tate 1898) [=*Strebloramphus mirulus]F27
Coptochetus asperulus [=Fusus asperulus]F27
Trifora Deshayes in Blainville 1828O27b
|--*T. perversaO27b
|--T. callipyrga Bartsch 1907O27b
|--T. carpenteri Bartsch 1903O27b
|--T. catalinensis Bartsch 1907O27b
|--T. montereyensis Bartsch 1907O27b
|--T. pedroana Bartsch 1907O27b
|--T. peninsularis Bartsch 1907O27b
`--T. stearnsi Bartsch 1907O27b
Barocospira Horný 1964FBL02
Ibergia Blodgett & Frýda 1999FBL02
|--Yukonoconcha Frýda, Blodgett & Lenz 2002FBL02
| `--*Y. pedderi Frýda, Blodgett & Lenz 2002FBL02
`--Crassimarginata Jhaveri 1969BR05
|--*C. crassicosta Jhaveri 1969FBL02
`--C. tenuicosta Jhaveri 1969FBL02
Levihelix pusilla Gründel 2000GK02
Adelphotectonica reeviHS01
Limulatys reliquusHS01
|--P. manawatawhiaHS01
|--P. olivaceaHS01
| |--P. o. olivaceaHS01
| `--P. o. impressaHS01
|--P. rekohuanaHS01
| |--P. r. rekohuanaHS01
| `--P. r. lactorubraHS01
|--P. semiplicataHS01
`--P. zosterophilaHS01
Camita rotellina (Gould 1860) [=Trochus (Monodonata) rotellinus]H08
|--T. maculata (Brazier 1877) [=Thalotia maculata]H09
|--T. monilifera Adams 1855H09
`--T. montrouzieri (Fischer 1878) [=Tectaria montrouzieri]H09
Mecoliotia spinosa Hedley 1902H09
Couthouyia aculeata Hedley 1900H09
|--O. tasmanica (Ten.-Woods 1877) [=Parthenia tasmanica]H09
`--O. terebellum [=Cingula terebellum; incl. O. exarata, Parthenia exarata]C64
Elusa subulata (Adams 1855) [=Pyramidella subulata]H09
Laimodonta conica Pease 1863K65
Paraplysia piperata (Smith 1884) [=Aplysia piperata]H09
Euselenops luniceps (Cuvier 1830) [=Pleurobranchus luniceps]H09
Placomopherus insignis Smith 1884H09
Adiozoptyxis Dietrich 1925 [Adiozoptyxinae, Adiozoptyxisinae]BR05
`--*A. polymorpha (Gemmellaro 1865) [=Nerinea polymorpha]BR17
‘Cavolinia’ Bruguière 1791 non Abilgaard 1791BR05
Crenea Risso 1826BR05
Cypraeogemmula Vredenburg 1920 [Cypraeogemmulinae, Cypraeogemmulini]BR05
`--*C. scabriuscula (Koenen 1890)BR17 [=Trivia scabriusculaBR17, Cypraea scabriusculaH09]
Incrispella Tasch 1963 [Incrispellidae]BR05
`--*I. rectotortis Tasch 1963BR17
Ladamarekia Horný 1992 [Ladamarekiidae]BR05
`--*L. miranda Horný 1992BR17
Melanioptyxis Cossmann 1896 [Melanioptyxinae]BR05
`--*M. altararis (Cossmann 1885) [=Nerinea altararis]BR17
Neoptyxis Wenz 1940 [Neoptyxidae]BR05
`--*N. astrachanica (Rebinder 1902) [=Nerinea astrachanica]BR17
Oligoptyxis Pchelintsev 1953 [Oligoptyxidae]BR05
`--*O. turricula Pchelintsev 1953BR17
Yuopisthonema Nützel in Bouchet, Rocroi et al. 2017BR17 (see below for synonymy)
`--*Y. undulatum (Yu 1974) [=*Opisthonema undulatum]BR17
Palaeocyclophorus Wenz 1923BR05
Pentaptyxis Pchelintsev 1965 [Pentaptyxidae]BR05
`--*P. staszycii (Zeuschner 1849) [=Acteon staszycii]BR17
Plesioplocus Pchelintsev 1953 [Plesioplocidae]BR05
`--*P. grandis Pchelintsev 1953BR17
Polyodonte Fischer 1807BR05
Polyodonta Megerle 1811BR05
Protoconchoididae [Protoconchioididae]BR05
|--Patelliconus Horný 1961BR17, BR05 [Patelliconidae]
| `--*P. primulus (Perner 1903) [=Palaeacmaea primula]BR17
`--Protoconchoides Shaw 1962 [=Protoconchioides (l. c.)]BR05
|--*P. hermitensis (Resser 1945) [=Scenella hermitensis]BR17
`--P. conulaTS11
Rhytidopilus Cossmann 1895 [=Rhytidophilus (l. c.); Rhytidophilidae, Rhytidopilidae]BR05
`--*R. humbertinus (Buvignier 1852) [=Patella humbertina]KC60
Stylocheilus Gould 1852 [Stylocheilinae, Stylochiniae]BR05
`--*S. lineolatus (Gould 1852) [=Aplysia lineolatus]BR17
Transovula de Gregorio 1880 [Transovulini]BR05
`--*T. schefferi (de Gregorio 1880) [=Ovula schefferi]BR17
Papustyla pulcherrimaUSDI77
Acilia franciscaeTE01
Bathyomphalus contortus (Linnaeus 1758)L06
Heterostoma Filippi 1837CC06, LT64 [=Heterostomum Diesing 1850LT64]
`--H. paupercula (Lowe 1831) [incl. Steenbergia duplex]CC06
|--S. depauperata (Lowe 1831) [incl. S. latinea (Paiva 1866)]CC06
|--S. obtectaCC06
`--S. squalidaCC06
Plagyrona placida (Shuttleworth 1852)CC06
Staurodon saxicolaCC06
|--A. arcinellaCC06
| |--A. a. arcinellaCC06
| `--A. a. papillosaCC06
|--A. crassiusculaCC06
|--A. effugiensCC06
|--A. morenensisCC06
`--A. promontoriensisCC06
|--P. loweiCC06
`--P. portosanctanaCC06
|--A. cimensisCC06
|--A. gracilisCC06
|--A. melampoidesCC06
|--A. oryzaCC06
|--A. terebellaCC06
|--A. triticeaCC06
`--A. tuberculataCC06
Cylichnidia ovuliformisCC06
Lemniscia michaudiCC06
Eufenella pupoides (Adams 1860)BD86
Clathrofenella reticulata (Adams 1860)BD86
Allocharopa belliSC07
|--P. gerassimoviZZ88
|--P. rammelmeyeriZZ88
`--P. vitimensisZZ88
Brotiopsis wakinoensisZZ88
Tulotomoides talatziensisZZ88
Gyrodema etheridgeiF71
Magelia bidens Tenison-Woods 1878F71
Odontostomia cretacea Etheridge 1907F71
Lindoliva spengleri Petuch 1988P89
Armiger crista (Linnaeus 1758)P88
Probythinella lacustris (Baker 1928)P88
Bulimnea megasoma (Say 1824)P88
|--J. ciliata (Sowerby 1843)AA88
`--J. stephanophora (Moellendorff 1895)AA88
Gymnentome validaH75
Ittibittium Houbrick 1993BC01
`--I. turriculum (Usticke 1969) (see below for synonymy)BC01
Torrella Henderson & Bartsch 1920BC01
`--*T. torreiana (Gundlach ex Arango y Molina 1878) [=Ctenopoma torreianum]BC01
Pseudotorinia retifera (Dall 1892) [incl. Heliacus crystallina Usticke 1969]BC01
Boonea impressa (Say 1822) [incl. Odostomia (Menestho) beauforti Jacot 1921]BC01
Fargoa bushiana (Bartsch 1909) [incl. Odostomia charlottae Usticke 1969]BC01
Aroapyrgus Baker 1931BC01
`--A. dubiosus (Adams 1852) [=Truncatella dubiosa]BC01
Chippewaella patellithecaW01
Moerkeia Böhm 1895NE02
Greggelix indigenaC90
Torquilla frumentumD56
Sinopupoides hubeiensisNG13
Bulgarica thessalonica (Olivier 1801)O03
Clavilithes rugosusC60
Partulida spiralisC60
Ampullella bulimoidesC60
Cepatia cepaceaC60
Palaeocyclotus exaratusC60
Pseudorhytidopilus Cox in Knight, Cox et al. 1960KC60
`--*P. lennieri Cox in Knight, Cox et al. 1960 (see below for synonymy)KC60
Atlantobellerophon Trechmann 1930KC60
`--*A. zealandicus Trechmann 1930KC60
Burdikinia Knight 1937 [=Polyamma Etheridge 1917 non Kriechbaumer 1894]KC60
`--*B. burdikiensis (Etheridge 1917) [=*Polyamma burdikiensis]KC60
Cinctaspira Powell 1933KC60
`--*C. conica Powell 1933KC60
‘Conchula’ Steininger 1849 non Herrmannsen 1847 (n. d.)KC60
`--*C. cylindracea Steininger 1849 (n. d.)KC60
‘Geinitzia’ Dietz 1911 nec Gemmellaro 1892 nec Handlirsch 1906KC60
`--*G. carinata Dietz 1911KC60
Kebina Vologdin 1955KC60
`--*K. pulchra Vologdin 1955KC60
Ozarkina Ulrich & Bridge 1931 [=Ozarkispira Ulrich & Bridge 1931 non Walcott 1924]KC60
`--*O. typica (Ulrich & Bridge 1931) [=*Ozarkispira typica]KC60
Ozarkocanus Heller 1956KC60
`--*O. prearcuatus Heller 1956KC60
Pondia Oder 1932KC60
`--*P. powelli Oder 1932KC60
Pythmenema Lamont & Gilbert 1945KC60
`--*P. praenantium (Phillips 1848) [=Euomphalus praenantius]KC60
Trochotremaria Ryckholt 1860KC60
Umbotrochus Perner 1903 [=Umbonitrochus Cossmann 1918]KC60
`--*U. aspersus Perner 1903KC60
Acevina Rusconi 1952KC60
`--*A. cuyunchensis (Rusconi 1952) [=Helcionella (*Acevina) cuyunchensis]KC60
|--O. abbotti Vanatta 1918V18
|--O. blandiV18
`--O. hilliV18
Aplysiopsis smithi (Marcus 1961)B77
Sorapisella Bandel 1993B00
Triadoskenea Bandel 1993B00
`--T. ampezzana Bandel 1993B02
Rolandomphalus Schwardt 1992B00
Macrophragmus tokyoensisK10
Fascioplex neozelanicaK95
Pseudoaclisina Yoo 1994B02
|--*P. turgida [=Aclisina turgida]B02
`--P. microspirulataB02
Palaeoalvania Yoo 1994B02
`--*P. talenti (Yoo 1994) [=Aclisina talenti]B02
Sinistriconcha Heidelberger & Bandel 1999B02
|--*S. lierli Heidelberger & Bandel 1999B02
`--S. minutissima (Yoo 1988)B02
|--I. dupuydelomeiE99
`--I. marmoratusC84
Scabrella Sacco 1890H22
`--*S. scabra [=Columbella scabra]H22
Pseudaulicina Chavan 1948BR17
`--*P. musicalis (Lamarck 1803) [=Voluta musicalis, Volutilithes musicalis]BR17
Kimina Yoo 1994B02
|--*K. globosa Yoo 1994B02
|--K. australis Yoo 1994B02
|--K. minor Yoo 1994B02
`--K. yooi Frýda & Bandel 1997B02
Harperispira Bandel 2002BR17
Kiringella Rozov 1968W01, RJ76
|--K. kultavasaensisRJ76
`--K. pyramidalisW01
Palaeoniso brazieriSS16
Stusakia pulchraF12
|--M. bajanohongorensisV-RR93
|--M. debilicostataV-RR93
`--M. shuwaloviV-RR93
Severynella trochlearis Hynda 1983D94
Lacunina Kittl 1899TTE93
Bullopsis Conrad 1858TTE93
Serradonta Okutani, Tsuchida & Fujikara 1992AJH07
Pleuroprocta silvaticaPB27
|--T. maculatumPB27
| |--T. m. maculatumPB27
| `--T. m. perforatumPB27
|--T. pardusPB27
`--T. ruwenzoriensePB27
|--B. germainiPB27
|--B. interiorisPB27
|--B. liocephalaPB27
`--B. runssorinaPB27
|--A. copiosusPB27
|--A. spatiosusPB27
`--A. tenebrosusPB27
|--L. apicalisE99
|--L. dufrenoyiE99
|--L. kleiniE99
|--L. leptolomaE99
|--L. pisumE99
`--L. valentinensisE99
|--L. v. valentinensisE99
`--L. v. sermenazensisE99
Wenzia ramondiE99
|--M. christoliE99
|--M. delphinensisE99
| |--M. d. delphinensisE99
| `--M. d. tersannensisE99
|--M. gualinoiE99
`--M. turonensisE99
Pseudoleacina (Paraglandina)E99
|--P. (P.) christolianaE99
`--P. (P.) oligostrophaE99
|--P. incrassatusE99
`--P. mollonensisE99
|--G. leobersdorfensisE99
`--G. locardiE99
|--N. bleicheriE99
|--N. raricostatusE99
|--N. suturalisE99
| |--N. s. suturalisE99
| `--N. s. gracilisE99
|--N. truciE99
`--N. villafranchianusE99
|--E. aloisiiE99
|--E. fischeriE99
`--E. patuliformisE99
|--M. castetiE99
`--M. collongeoniE99
Fortuna seringiE99
Truciella ballesioiE99
|--P. bernardiiE99
|--P. nordsieckiE99
`--P. plioauriculatumE99
Monoptychia monoptyxE99
Serruluna anodonE99
Bellerophontacea [Belleromorpha, Bellerophontina, Fissidorsata, Planspiralia, Prorhipidoglossa]KC60
| i. s.: Patellostium Waagen 1880 (n. d.)KC60
| `--*P. macrostoma (Roemer 1844) [=Bellerophon macrostoma]KC60
| Euphemitella Tasch 1953 (n. d.)KC60
| `--*E. emrichi Tasch 1953KC60
`--Isospira Koken 1897KC60 [Isospiridae, IsospiroideaBR05]
`--*I. bucanioides Koken 1897BR17
Utriculopsis globosa (Lovén 1846) (see below for synonymy)N79
Patelloidea Tasch 1961T61
`--*P. limnensis Tasch 1961T61
Permoplanorboidea Tasch 1961T61
`--*P. primus Tasch 1961T61
Rugoplanorboidella Tasch 1961T61
`--*R. sedgwickii Tasch 1961T61
Wellingtonia Tasch 1961T61
`--*W. producta Tasch 1961T61
Tretospira divesuralicaG31
Sosiolites vassilievkensisG31
|--P. inornataG31
`--P. nitidulumG31
Isognomostoma isognomostomaJ63
Platyla politaNB19
|--P. p. politaNB19
`--P. p. regina (Subai 1977)NB19
Paleobulimulus eocenicus Parodiz 1949P63
Sigatica semisulcata (Gray in Beechey 1839) [=Natica semisulcata]MP65
Hemitrochus variansJ65
|--P. gregorianaJ65
`--P. macrodonJ65
|--P. m. macrodonJ65
`--P. m. utowanaJ65
Bigotella Cossmann 1914EH19
Vertagus graniferus Pease 1861 [=Cerithium graniferum]K65
|--S. bipes Pease 1860K65
|--S. elongata Pease 1860K65
`--S. grandis Pease 1860K65
|--P. bellum Pease 1860K65
`--P. ornatum Pease 1860K65
Histiophorous maculatus Pease 1860K65
Dentiora rubida Pease 1863K65
Mitroidea multiplicata Pease 1865K65
|--R. sebaeF66
`--R. umbilicataP79 [incl. R. globosa Tenison Woods 1876P79, TW76]
|--C. aureusTW76
`--C. ornatus Tenison Woods 1876TW76
Discotectonica wannonensisBRW98
Nanamoria strophonBRW98
Leiostomus Swainson 1840 [incl. Sycostoma Cox 1931]BS00
`--*Sycostoma’ bulbiformeBS00
Mirochiliticus mirusX89
Singulitubus obliquusX89
Guizhouspira convexaX89
Turbinilopsis rotundusX89
Euthystylus costatusX89

Acanthonema newberryi (Meek 1871)TTE93 [=Orthonema newberryiKC60; incl. *A. holopiforme Grabau 1909KC60, BR17]

Acteonina d’Orbigny 1850BR05 [=Actaeonina (l. c.)BR05; Acteoninidae, Acteonininae, ActeoninoideaBR17]

Apogastropoda [Anaclodonta, Architectonicoida, Chalazaeata, Ctenobranchia, Davisianoidei, Dioecia, Discopoda, Ectobranchia, Entobranchia, Entomotaeniata, Gymnoglossa, Heterogastropoda, Heterostropha, Holonephridia, Homoestropha, Hydrocenoidei, Iniophthalma, Loxonematacea, Monopleurobranchia, Monotocardia, Murchisoniata, Musioglossata, Oligopteria, Orbacea, Paludinimorpha, Pectinibranchia, Pectinibranchiata, Peristomacea, Peristomania, Peristomidae, Pneumonobranchiata, Potamophila, Proboscidifera, Procaenogastropoda, Ptenobranchiata, Pyramidellida, Pyramidellimorpha, Pyramidelliones, Pyramidelloida, Rissoellina, Solenisciformes, Spartaebranchia, Subulitiformes, Taenioglossa, Trachelobranchia, Volumina]S11

Caenogastropoda [Architaenioglossa, Architaenioglossae, Basiophthalma, Cyclophoracea, Cyclophoroidei, Fusiformia, Globularioidei, Mesogastropoda, Mesogastropodida, Opisthophthalma, Palaeocyclophoridae, Procyclophoridae, Procyclophoroidea, Rostrifera, Teleogeophila]S11

Dendropupa Owen 1859BR17 [incl. Strophella Dawson 1895KC60, Strophites Dawson 1880 non Deshayes 1832KC60; Dendropupidae]

Gastropoda [Abranchiata, Alata, Anentomostomata, Anisobranchia, Anisopleura, Aponotoneura, Apterygia, Archaeogastropoda, Archaeoprosobranchia, Arthrocochlides, Asiphonobranchia, Asiphonata, Aspidobranchia, Auxogastropoda, Cervicobranchiata, Chiastoneura, Chismobranchiata, Clypeaceae, Cocculinacea, Cocculiniformia, Cochliostraca, Conivalvia, Cricostomata, Ctenobranchiata, Cyclobranchia, Dicoeca, Dicranobranchia, Diotocardia, Eogastropoda, Fissobranchia, Fissuracea, Fornices, Galeroconcha, Gasteropoda, Gasteropodophora, Gasterozoa, Gastraeopoda, Gastropodea, Goniostomata, Gonyostomata, Gymnopoda, Helcionellina, Hemicyclostoma, Holostomata, Hyperstrophina, Lepadophora, Lepetellina, Limaces, Neurobranchia, Operculata, Orthogastropoda, Otidea, Paracephalophora, Paragastropoda, Patellina, Peristomatacea, Peristomatidae, Platyceratacea, Podophthalma, Prosobranchia, Prosobranchiata, Pselaphocephala, Raphidoglossa, Rhipidoglossa, Riphidoglossa, Scutiformia, Sinuata, Sinuopeoidei, Solenostomata, Streptoneura, Syncephala, Tectipeda, Triforidoidei, Trochacea, Trochina, Trochomorpha, Trochomorphi, Turbinacea, Turbinimorpha, Xenophoroidei]

Ittibittium turriculum (Usticke 1969) [=Bittium varium f. turricula, Alabina turricula, B. turriculum, Cerithiodes turricula]BC01

Kittlidiscus Haas 1953BR05 [=Schizodiscus Kittl 1891 non Hall & Clarke 1888KC60; KittlidiscidaeBR17]

*Kittlidiscus bronni (Klipstein 1845)KC60 [=Pleurotomaria bronniKC60, Schizodiscus bronniKC60; incl. P. plana Klipstein 1845 non Münster 1844KC60, *Kittlidiscus planusBR17, *Schizodiscus planusBR17]

*Pseudorhytidopilus lennieri Cox in Knight, Cox et al. 1960 [incl. Helcion castellana Lennier 1868 non Patella castellana Thurmann & Étallon 1861]KC60

Sorbeoconcha [Anandria, Aulobranchiata, Brevicommisurata, Canalifera, Eriophthalma, Neotaenioglossa, Omalaxoidea, Palaeocaenogastropoda, Purpurifera, Siphonobranchei, Siphonobranchia, Siphonobranchiata, Tubispiracea, Tubispirantia]S11

Spirostylus linctus (Böhm 1895) [=Omphaloptycha lincta; incl. O. porrecta Böhm 1895]TE01

Utriculopsis globosa (Lovén 1846) [=Amphisphyra globosa, Utriculus globosus; incl. Utriculopsis vitrea Sars 1870]N79

Yuopisthonema Nützel in Bouchet, Rocroi et al. 2017BR17 [=Opisthonema Yu 1974 non Gill 1862BR05; Opisthonematidae, Yuopisthonematidae]

*Type species of generic name indicated


[AJH07] Amano, K., R. G. Jenkins & Y. Hikida. 2007. A new gigantic Nucinella (Bivalvia: Solemyoida) from the Cretaceous cold-seep deposit in Hokkaido, northern Japan. Veliger 49 (2): 84–90.

[AA88] Auffenberg, K., & T. Auffenberg. 1988. Density, spatial distribution, activity patterns, and biomass of the land snail, Geophorus bothropoma Moellendorff (Prosobranchia: Helicinidae). Nautilus 102 (1): 40–45.

[B63] Baker, H. B. 1963. Paludomidae (Pleuroceridae). Nautilus 77 (1): 34–35.

[B00] Bandel, K. 2000. The new family Cortinellidae (Gastropoda, Mollusca) connected to a review of the evolutionary history of the subclass Neritimorpha. Neues Jahrbuch für Geologie und Paläontologie—Abhandlungen 217 (1): 111–129.

[B02] Bandel, K. 2002. About Heterostropha (Gastropoda) from the Carboniferous and Permian. Mitteilungen des Geologisch-Paläontologischen Institutes der Universität Hamburg 86: 45–80.

[BS00] Bandel, K., & W. Stinnesbeck. 2000. Gastropods of the Quiriquina Formation (Maastrichtian) in central Chile: paleobiogeographic relationships and the description of a few new taxa. Zbl. Geol. Paläont. Teil I 7–8: 757–788.

[BD86] Barash, A., & Z. Danin. 1986. Further additions to the knowledge of Indo-Pacific Mollusca in the Mediterranean Sea (Lessepsian migrants). Spixiana 9 (2): 117–141.

[BRW98] Beesley, P. L., G. J. B. Ross & A. Wells (eds) 1998. Fauna of Australia vol. 5. Mollusca: The Southern Synthesis. Part A. Australian Biological Resources Study: Canberra.

[B77] Bertsch, H. 1977. The Chromodoridinae nudibranchs from the Pacific coast of America.—Part I. Investigative methods and supra-specific taxonomy. Veliger 20 (2): 107–118.

[B96] Bond, C. E. 1996. Biology of Fishes 2nd ed. Saunders College Publishing: Fort Worth.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[C84] Cain, A. J. 1984. Heterosematism in snails. Malacologia 25 (1): 161–172.

[CC06] Cameron, R. A. D., L. M. Cook, G. A. Goodfriend & M. B. Seddon. 2006. Fossil land snails of Porto Santo, Madeiran archipelago: change and stasis in Pleistocene to Recent times. Malacologia 49 (1): 25–59.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[C60] Cox, L. R. 1960. Gastropoda: general characteristics of Gastropoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I84–I169. Geological Society of America, and University of Kansas Press.

[C90] Crawford, C. S. 1990. Scorpiones, Solifugae, and associated desert taxa. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 421–475. John Wiley & Sones: New York.

[D56] Dawes, B. 1956. The Trematoda with special reference to British and other European forms. University Press: Cambridge.

[D94] Dzik, J. 1994. Evolution of ‘small shelly fossils’ assemblages of the early Paleozoic. Acta Palaeontologica Polonica 39 (3): 247–313.

[EH19] Énay, R., & M. K. Howarth. 2019. Part L, revised, volume 3B, chapter 7: Systematic descriptions of the Perisphinctoidea. Treatise Online 120: 1–184.

[E99] Esu, D. 1999. Contribution to the knowledge of Neogene climate changes in western and central Europe by means of non-marine molluscs. In: J. Agustí, L. Rook & P. Andrews (eds) The Evolution of Neogene Terrestrial Ecosystems in Europe pp. 328–354. Cambridge University Press.

[F26] Finlay, H. J. 1926. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F27] Finlay, H. J. 1927. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488-533.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[F66] Fraser, L. 1866. Communication of a list of mollusks collected by R. Swinhoe, Esq., in Formosa. Proceedings of the Zoological Society of London 1866: 146.

[F01] Frýda, J. 2001. Discovery of a larval shell in Middle Paleozoic subulitoidean gastropods with description of two new species from the Early Devonian of Bohemia. Vĕstnik Českého Geologického Ústavu 76 (1): 29–37.

[F12] Frýda, J. 2012. Phylogeny of Palaeozoic gastropods inferred from their ontogeny. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 395–435. Springer.

[FB01] Frýda, J., & R. B. Blodgett. 2001. The oldest known heterobranch gastropod, Kuskokwimia gen. nov., from the Early Devonian of west-central Alaska, with notes on the early phylogeny of higher gastropods. Vĕstnik Českého Geologického Ústavu 76 (1): 39–53.

[FB04] Frýda, J., & R. B. Blodgett. 2004. New Emsian (Late Early Devonian) gastropods from Limestone Mountain, Medfra B-4 quadrangle, west-central Alaska (Farewell Terrane), and their paleobiogeographic affinities and evolutionary significance. Journal of Paleontology 78 (1): 111–132.

[FBL02] Frýda, J., R. B. Blodgett & A. C. Lenz. 2002. New Early Devonian gastropods from the families Crassimarginatidae (new family) and Scoliostomatidae (new family), Royal Creek area, Yukon Territory, Canada. Journal of Paleontology 76 (2): 246–255.

[GO06] Giribet, G., A. Okusu, A. R. Lindgren, S. W. Huff, M. Schrödl & M. K. Nishiguchi. 2006. Evidence for a clade composed of molluscs with serially repeated structures: monoplacophorans are related to chitons. Proceedings of the National Academy of Sciences of the USA 103 (20): 7723–7728.

[GS75] Golikov, A. N., & Y. I. Starobogatov. 1975. Systematics of prosobranch gastropods. Malacologia 15 (1): 185–232.

[G31] Grabau, A. W. 1931. The Permian of Mongolia: A report on the Permian fauna of the Jisu Honguer limestone of Mongolia and its relations to the Permian of other parts of the world. American Museum of Natural History: New York.

[G88] Gray, J. 1988. Evolution of the freshwater ecosystem: the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology 62: 1–214.

[GK02] Gründel, J., & T. Kowalke. 2002. Palaeorissoinidae, a new family of marine and brackish water Rissooidea (Gastropoda, Littorinimorpha). Neues Jahrbuch für Geologie und Paläontologie Abhandlungen 226 (1): 43–57.

[GAS03] Gulbin, V. V., I. S. Arzamastsev & V. M. Shulkin. 2003. Ecological monitoring of the water area of Port Vostochnyi (Wrangel Bay) in the Sea of Japan (1995–2002). Russian Journal of Marine Biology 29 (5): 284–295.

[H75] Häntzschel, W. 1975. Treatise on Invertebrate Paleontology pt W. Miscellanea Suppl. 1. Trace Fossils and Problematica 2nd ed. The Geological Society of America: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[HS01] Hayward, B. W., A. B. Stephenson, M. S. Morley, W. M. Blom, H. R. Grenfell, F. J. Brook, J. L. Riley, F. Thompson & J. J. Hayward. 2001. Marine biota of Parengarenga Harbour, Northland, New Zealand. Records of the Auckland Museum 37: 45–80.

[H08] Hedley, C. 1908. Studies on Australian Mollusca. Part X. Proceedings of the Linnean Society of New South Wales 33: 456–489, pls 7–10.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H22] Hedley, C. 1922. A revision of the Australian Turridae. Records of the Australian Museum 13 (6): 213–359, pls 42–56.

[HP04] Herbert, G. S., & R. W. Portell. 2004. First paleontological record of larval brooding in the calyptraeid gastropod genus Crepidula Lamarck, 1799. Journal of Paleontology 78 (2): 424–429.

[J65] Jacobson, M. K. 1965. On some land shells of Eleuthera, Bahamas. Nautilus 78 (4): 120–125.

[J63] Jánossy, D. 1963. Letztinterglaziale Vertebraten-Fauna aus der Kálmán Lambrecht-Höhle (Bükk-Gebirge, nordost-Ungarn) I. Acta Zoologica Academiae Scientiarum Hungaricae 9 (3–4): 293–331.

[JB12] Johnson, M. E., & B. G. Baarli. 2012. Development of intertidal biotas through Phanerozoic time. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 63–128. Springer.

[KBC03] Kashin, I. A., E. V. Bagaveeva & S. F. Chaplygina. 2003. Fouling communities of hydrotechnical constructions in Nakhodka Bay (Sea of Japan). Russian Journal of Marine Biology 29: 267–283.

[K65] Kay, E. A. 1965. Marine molluscs in the Cuming collection, British Museum (Natural History) described by William Harper Pease. Bulletin of the British Museum (Natural History): Zoology Supplement 1: 1–96, 14 pls.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[K95] Köhler, R. 1995. A new species of the fossil turtle Psephophorus (order Testudines) from the Eocene of the South Island, New Zealand. Journal of the Royal Society of New Zealand 25: 371–384.

[K10] Kostina, E. E. 2010. Distribution of macrobenthic communities in the intertidal zone of the Vietnamese islands. In: China-Russia Bilateral Symposium: Proceedings of the China-Russia Bilateral Symposium of “Comparison on Marine Biodiversity in the Northwest Pacific Ocean”, 10–11 October 2010, Qingdao (China) pp. 109–111. Institute of Oceanology, Chinese Academy of Sciences; A. V. Zhirmunsky Institute of Marine Biology, Far East Branch of the Russian Academy of Sciences.

[L06] Lewin, I. 2006. The gastropod communities in the lowland rivers of agricultural areas—their biodiversity and bioindicative value in the Ciechanowska Upland, central Poland. Malacologia 49 (1): 7–23.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 2. The Geological Society of America and The University of Kansas Press.

[MP65] Merrill, A. S., & R. E. Petit. 1965. Mollusks new to South Carolina. Nautilus 79 (2): 58–66.

[M03] Meyer, C. P. 2003. Molecular systematics of cowries (Gastropoda: Cypraeidae) and diversification patterns in the tropics. Biological Journal of the Linnean Society 79: 401–459.

[M62] Monniot, F. 1962. Recherches sur les graviers a Amphioxus de la région de Banyuls-sur-Mer. Vie et Milieu 13: 231–322.

[NB19] Nardi, G., A. Braccia, S. Cianfanelli & M. Bodon. 2019. Revision of the systematic position of Lindbergia garganoensis Gittenberger & Eikenboom, 2006, with reassignment to Vitrea Fitzinger, 1833 (Gastropoda, Eupulmonata, Pristilomatidae). Basteria 83 (1–3): 19–28.

[NG13] Ni, X., D. L. Gebo, M. Dagosto, J. Meng, P. Tafforeau, J. J. Flynn & K. C. Beard. 2013. The oldest known primate skeleton and early haplorhine evolution. Nature 498: 60–64.

[N79] Norman, A. M. 1879. The Mollusca of the fiords near Bergen, Norway. Journal of Conchology 2: 8–77.

[NE02] Nützel, A., & D. H. Erwin. 2002. Battenizyga, a new Early Triassic gastropod genus with a discussion of the caenogastropod evolution at the Permian/Triassic boundary. Paläontologische Zeitschrift 76 (1): 21–27.

[OT04] Olabarria, C., & M. H. Thurston. 2004. Patterns of morphological variation of the deep-sea gastropod Troschelia berniciensis (King, 1846) (Buccinidae) from the northeastern Atlantic Ocean. Journal of Molluscan Studies 70: 59–66.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. 2 pt 2. Stanford University Press: Stanford University (California).

[O03] Örstan, A. 2003. The rediscovery of Zonites algirus in İstanbul, Turkey (Gastropoda: Pulmonata: Zonitidae). Zoology in the Middle East 29: 75–78.

[PE02] Pan, H.-Z., & D. H. Erwin. 2002. Gastropods from the Permian of Guangxi and Yunnan Provinces, south China. Journal of Paleontology 76 (Memoir 56): 1–49.

[P63] Parodiz, J. J. 1963. New fresh-water Mollusca from the Eogene of Chile and Patagonia. Nautilus 76 (4): 145–148.

[P79] Petterd, W. F. 1879. Critical remarks on the Rev. J. E. Tenison Woods’ “Census of Tasmanian shells”. Journal of Conchology 2: 353–354.

[P89] Petuch, E. J. 1989. New species of Malea (Gastropoda: Tonnidae) from the Pleistocene of southern Florida. Nautilus 103 (3): 92–95.

[P02] Pickford, M. 2002. Early Miocene grassland ecosystem at Bukwa, Mount Elgon, Uganda. Comptes Rendus Palevol 1: 213–219.

[PB27] Pilsbry, H. A., & J. Bequaert. 1927. The aquatic mollusks of the Belgian Congo, with a geographical and ecological account of Congo malacology. Bulletin of the American Museum of Natural History 53 (2): 69–602, pls 10–77.

[P88] Pip, E. 1988. Niche congruency of freshwater gastropods in central North America with respect to six water chemistry parameters. Nautilus 102 (2): 65–72.

[PC08] Ponder, W. F., D. J. Colgan, J. M. Healy, A. Nützel, L. R. L. Simone & E. E. Strong. 2008. Caenogastropoda. In: Ponder, W. F., & D. L. Lindberg (eds) Phylogeny and Evolution of the Mollusca pp. 331–383. University of California Press: Berkeley.

[R42] Reeve, L. 1842. Descriptions of two new species of Oniscia, a genus of pectinibranchiate mollusks. Proceedings of the Zoological Society of London 10: 91.

[RJ76] Runnegar, B., & P. A. Jell. 1976. Australian Middle Cambrian molluscs and their bearing on early molluscan evolution. Alcheringa 1 (2): 109–138.

[S11] Simone, L. R. L. 2011. Phylogeny of the Caenogastropoda (Mollusca), based on comparative morphology. Arquivos de Zoologia 42 (4): 161–323.

[SC07] Stanisic, J., R. A. D. Cameron, B. M. Pokryszko & J. C. Nekola. 2007. Forest snail faunas from S. E. Queensland and N. E. New South Wales (Australia): patterns of local and regional richness and differentiation. Malacologia 49 (2): 445–462.

[SS16] Süssmilch, C. A., & W. G. Stone. 1916. Geology of the Jenolan Caves district. Journal and Proceedings of the Royal Society of New South Wales 49 (3): 332–384, pls 55–56.

[T61] Tasch, P. 1961. Paleolimnology: Part 2—Harvey and Sedgwick counties, Kansas: stratigraphy and biota. Journal of Paleontology 35: 836–865.

[TW76] Tenison Woods, J. E. 1876. Description of new Tasmanian shells. Papers and Proceedings of the Royal Society of Tasmania 1875: 134–162.

[TRM20] Thomas, R. D. K., B. Runnegar & K. Matt. 2020. Pelagiella exigua, an early Cambrian stem gastropod with chaetae: lophotrochozoan heritage and conchiferan novelty. Palaeontology 63 (4): 601–627.

[TE01] Tong, J., & D. H. Erwin. 2001. Triassic gastropods of the southern Qinling Mountains, China. Smithsonian Contributions to Paleobiology 92: 1–47.

[TS11] Tortello, M. F., & N. M. Sabattini. 2011. Totoralia, a new conical-shaped mollusk from the Middle Cambrian of western Argentina. Geologica Acta 9 (2): 175–185.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[USDI77] USDI (United States Department of the Interior). 1977. Endangered and threatened wildlife and plants—republication of list of species. Federal Register 42: 36420–36431.

[V18] Vanatta, E. G. 1918. Two new shells from Hayti. Proceedings of the Academy of Natural Sciences of Philadelphia 70: 237–238.

[V-RR93] Vickers-Rich, P., & T. H. Rich. 1993. The ICI Australia Catalogue of the Great Russian Dinosaurs Exhibition 1993–1995. Monash University.

[W01] Wagner, P. J. 2001. Gastropod phylogenetics: progress, problems and implications. Journal of Paleontology 75 (6): 1128–1140.

[X89] Xi Y.-H. 1989. Late Permian–Early Triassic Gastropods from Zhongying, Qinglong of Guizhou with analysis of their feeding habits and forms. Acta Palaeontologica Sinica 28 (5): 683–686.

[ZZ88] Zhu G.-X. & Zhou Y.-Z. 1988. Fresh-water molluscan fauna across Jurassic-Cretaceous boundary in southeast Jilin. Acta Palaeontologica Sinica 27 (6): 713–721.

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