Longitudinal section of Pachyphloia ovata under plane-polarised light, from Groves et al. (2004).

Belongs within: Nodosariina.

The Geinitzinidae are a group of uniserial foraminifera known from the Late Carboniferous and Permian, with an ancestrally atelo-monolamellar (i.e. lacking secondary lamellarity), hyaline-radial wall structure. In the derived genus Pachyphloia, the wall becomes plesio-monolamellar, with secondary lamellae enveloping some but not all previous chambers (Groves et al. 2003).

Geinitzinidae [Geinitzinacea]
| i. s.: Omoloniella Karavaeva & Pronina 2007M13
| Frondilina Bykova 1952HW93
| `--*F. devexis Bykova 1952LT64
|--Vervilleina Groves in Groves & Boardman 1999GAR03
| `--V. bradyi (Spandel 1901)GAR03
|--Nodosinelloides Mamet & Pinard 1992GAR03
| |--N. netschajewi (Cherdyntsev 1914)GAR03
| |--N. pinardae Groves & Wahlmann 1997GRA04
| `--N. ronda (Lipina 1949)GAR03
`--+--Geinitzina Spandel 1901GAR03 [=Geinitzella Spandel 1898 non Waagen & Wentzel 1866LT64]
| |--‘Textularia’ jonesi Brady 1876 (see below for synonymy)LT64
| |--G. ciscoensisC40
| |--G. postcarbonica Spandel 1901GAR03 [incl. G. triangularisF71]
| |--G. tauricaFP12
| `--G. tcherdynzevi Miklukho-Maclay 1954W76
`--+--Howchinella Palmieri in Foster et al. 1985GAR03
| |--*H. woodwardi (Howchin 1895) [=Frondicularia woodwardi]GAR03
| `--‘Geinitzina’ caseyi Crespin 1958GAR03
|--Maichelina Sosnina 1977HW93
|--Robustopachyphloia Lin 1980HW93
`--Pachyphloia Lange 1925GAR03 (see below for synonymy)
|--*P. ovata Lange 1925LT64
|--P. asymmetrica (Miklukho-Maklay 1954) [=*Parapachyphloia asymmetrica]LT64
|--P. cukurkoyi Sellier de Civrieux & Dessauvagie 1965GRA04
|--P. depressa (Miklukho-Maklay 1954) [=*Parageinitzina depressa]LT64
|--P. gefoensis (Miklukho-Maklay 1953) [=*Parapermodiscus gefoensis]LT64
|--P. magna (Miklukho-Maklay 1954) [=*Pseudogeinitzina magna]LT64
`--P. pigmobesa Wang 1976W76

Pachyphloia Lange 1925GAR03 [incl. Parageinitzina Miklukho-Maklay 1954LT64, Parapachyphloia Miklukho-Maklay 1954LT64, Parapermodiscus Miklukho-Maklay 1953LT64, Pseudogeinitzina Miklukho-Maklay 1954LT64; Pachyphloiidae]

‘Textularia’ jonesi Brady 1876 [=Lunucammina jonesi; incl. Textularia cuneiformis Jones in King 1850 non d’Orbigny 1876, *Geinitzella cuneiformis, *Geinitzina cuneiformis]LT64

*Type species of generic name indicated


[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use (3rd ed.) Harvard University Press: Cambridge (Massachusetts).

[FP12] Farabegoli, E., & M. C. Perri. 2012. Millennial physical events and the end-Permian mass mortality in the western Palaeotethys: timing and primary causes. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 719–758. Springer.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[GAR03] Groves, J. R., D. Altiner & R. Rettori. 2003. Origin and early evolutionary radiation of the order Lagenida (Foraminifera). Journal of Paleontology 77 (5): 831–843.

[GRA04] Groves, J. R., R. Rettori & D. Altiner. 2004. Wall structures in selected Paleozoic lagenide Foraminifera. Journal of Paleontology 78: 245–256.

[HW93] Hart, M. B., & C. L. Williams. 1993. Protozoa. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 43–70. Chapman & Hall: London.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt. C. Protista 2, vol. 1. The Geological Society of America and The University of Kansas Press.

[M13] Mikhalevich, V. I. 2013. New insight into the systematics and evolution of the Foraminifera. Micropaleontology 59 (6): 493–527.

[W76] Wang K. 1976. The foraminifera from the Changhsing Formation in western Guizhou. Acta Palaeontologica Sinica 15 (2): 187–195.

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