Eyebar goby Gnatholepis anjerensis, copyright Rickard Zerpe.

Belongs within: Gobiomorpharia.
Contains: Sicydiinae, Stenogobius, Ctenogobius, Periophthalmidae, Gobioididae.

The Gobionellidae are a clade of gobies with five branchiostegals and fused pelvic fins, primarily supported by molecular data (Thacker 2009).

    |  i. s.: Gobionellus [Gobionellinae]T09
    |           `--G. shufeldtiB96
    |         Lepidogobius lepidusND13
    |         Hazeus Jordan & Snyder 1901H27
    |           `--H. baliuris (Cuvier & Valenciennes 1837)GM90
    |--+--Acanthogobius Gill 1859T09, H27
    |  |    |--A. flavimanusT09
    |  |    |--A. lactipesT90
    |  |    `--A. ommaturus (Richardson 1845) [=Gobius ommaturus]H27
    |  `--+--Gobiopterus semivestitusT09
    |     `--Pandaka Herre 1927T09, S59
    |          |--*P. pusilla Herre 1927S59
    |          |--P. lidwilliT09
    |          |--P. minuta Smith 1959S59
    |          `--P. pygmaea Herre 1927S59
       |  |    |--M. abeiT90
       |  |    |--M. fusculus (Nichols 1951) [=Gobius (Tamanka) fusculus]AC90
       |  |    `--M. rivulusT09
       |  `--+--+--Eucyclogobius newberryiT09
       |     |  `--+--Gillichthys mirabilisT09
       |     |     `--Typhlogobius Steindachner 1879T09, TP86
       |     |          `--T. californiensis Steindachner 1879TP86
       |     `--ChaenogobiusT09
       |          |--C. annularisT09
       |          |--C. castaneusI92
       |          |--C. isazaT90
       |          |--C. laevisI92
       |          |--C. taranetziI92
       |          `--C. urotaeniaT90
          |  |  |  `--Awaous Steindachner 1860T09, H27
          |  |  |       |--A. guamensisT09
          |  |  |       |--A. ocellaris (Broussonet 1782)M58 (see below for synonymy)
          |  |  |       `--A. taiasicaF16
          |  |  `--StenogobiusT09
          |  `--+--+--CtenogobiusT09
          |     |  `--EvorthodusT09
          |     |       |--E. breviceps Gill 1859F16
          |     |       `--E. minutusT09
          |     `--Gnatholepis Bleeker 1874T09, S59
          |          |  i. s.: G. baliurus (Valenciennes 1837)S59
          |          |         G. calliurus Jordan & Seale 1905S59
          |          |         G. davaoensis Seale 1909H27
          |          |         G. deltoides (Seale 1901) [=Gobius deltoides]H27
          |          |         G. gemmeus Herre 1927H27
          |          |         G. inconsequens Whitley 1958H90
          |          |         G. volcanus Herre 1927H27
          |          |--*G. anjerensis (Bleeker 1850)S59, T09, S59 [=Gobius anjerensisS59]
          |          `--+--G. scapulostigmaT09
          |             `--G. thompsoniT09
               |  i. s.: OxudercesB96
                  |  `--PseudapocryptesT09
                  |       |--P. borneensisJR10
                  |       |--P. elongatusT09
                  |       `--P. lanceolatusAC90
                       |--Apocryptes glyphisodonH27
                       |--Scartelaos Swainson 1839T09, H27 [incl. BoleopsJR10]
                       |    |--S. histophorusT09
                       |    `--S. viridis (Hamilton-Buchanan 1822)M58 (see below for synonymy)
                       |--Boleophthalmus Valenciennes in Cuvier & Valenciennes 1837M58, H27
                       |    |--B. boddarti (Pallas 1770) (see below for synonymy)M58
                       |    `--B. chinensis (Osbeck 1771) (see below for synonymy)H27
                       `--Apocryptodon Bleeker 1874M58, H27
                            |--*A. madurensisH27
                            |--A. montalbani Herre 1927H27
                            |--A. sealei Herre 1927H27
                            |--A. taylori Herre 1927H27
                            `--A. wirzi Koumans 1837M58

Awaous ocellaris (Broussonet 1782)M58 [=Gobius ocellarisM58, Chonophorus ocellarisH27; incl. Awaous crassilabris Seale 1906M58]

Boleophthalmus boddarti (Pallas 1770) [=Gobius boddarti; incl. B. dentatus Weber 1908, B. sculptus Weber 1908]M58

Boleophthalmus chinensis (Osbeck 1771) [=Apocryptes chinensis; incl. Gobius pectinirostris Gmelin 1788, Boleophthalmus pectinirostris]H27

Scartelaos viridis (Hamilton-Buchanan 1822)M58 [=Gobius viridisM58, Boleophthalmus viridisM58; incl. B. aucupatorius Richardson 1845H27, B. novaeguineae Hase 1914M58]

*Type species of generic name indicated


[AC90] Allen, G. R., & D. Coates. 1990. An ichthyological survey of the Sepik River, Papua New Guinea. Records of the Western Australian Museum Supplement 34: 31–116.

[B96] Bond, C. E. 1996. Biology of Fishes 2nd ed. Saunders College Publishing: Fort Worth.

[F16] Fowler, H. W. 1916. The fishes of Trinidad, Grenada, and St. Lucia, British West Indies. Proceedings of the Academy of Natural Sciences of Philadelphia 67 (3): 520–548.

[GM90] Gill, H. S., & P. J. Miller. 1990. A new genus and species of goby from the Swan-Avon Estuary, Western Australia, with a redescription of the genus Favonigobius Whitley, 1930. Records of the Western Australian Museum 14 (4): 503–525.

[H27] Herre, A. W. 1927. Gobies of the Philippines and the China Sea. Bureau of Printing: Manila.

[H90] Hutchins, J. B. 1990. Fish survey of South Passage, Shark Bay, Western Australia. In: Berry, P. F., S. D. Bradshaw & B. R. Wilson (eds) Research in Shark Bay: Report of the France-Australe Bicentenary Expedition Committee pp. 263–278. Western Australian Museum.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[JR10] Jordan, D. S., & R. E. Richardson. 1910. Check-list of the species of fishes known from the Philippine archipelago. Bureau of Printing: Manila.

[M58] Munro, I. S. R. 1958. The fishes of the New Guinea region: a check-list of the fishes of New Guinea incorporating records of species collected by the Fisheries Survey Vessel “Fairwind” during the years 1948 to 1950. Papua and New Guinea Agricultural Journal 10 (4): 97–369 (reprinted: 1958. Territory of Papua and New Guinea Fisheries Bulletin no. 1).

[ND13] Near, T. J., A. Dornburg, R. I. Eytan, B. P. Keck, W. L. Smith, K. L. Kuhn, J. A. Moore, S. A. Price, F. T. Burbrink, M. Friedman & P. C. Wainwright. 2013. Phylogeny and tempo of diversification in the superradiation of spiny-rayed fishes. Proceedings of the National Academy of Sciences of the USA 110 (31): 12738–12743.

[S59] Smith, J. L. B. 1959. Gobioid fishes of the families Gobiidae, Periophthalmidae, Trypauchenidae, Taenioididae and Kraemeriidae of the western Indian Ocean. Ichthyological Bulletin 13: 185–225.

[T90] Taguchi, S. 1990. Nihon no Sakana. Kogakukan: Tokyo.

[T09] Thacker, C. E. 2009. Phylogeny of Gobioidei and placement within Acanthomorpha, with a new classification and investigation of diversification and character evolution. Copeia 2009 (1): 93–104.

[TP86] Thinès, G., & G. Proudlove. 1986. Pisces. In: Botosaneanu, L. (ed.) Stygofauna Mundi: A Faunistic, Distributional, and Ecological Synthesis of the World Fauna inhabiting Subterranean Waters (including the Marine Interstitial) pp. 709–733. E. J. Brill/Dr W. Backhuys: Leiden.

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