Gomphoidea

Mating pair of yellow-striped hunters Austrogomphus guerini, copyright Fir0002/Flagstaffotos.

Belongs within: Eteoanisoptera.

The Gomphoidea are a group of mostly small dragonflies, in which the larvae have a flattened prementum lacking a median cleft. The Late Jurassic and Early Cretaceous Proterogomphidae probably represent early members of this clade. The majority of living species are included in the family Gomphidae except the larger palaeotropical genus Ictinogomphus in the Lindeniidae, distinguished by the presence of crossveins in the triangle of the fore- and hind wings (Theischinger & Hawking 2006). Within the Gomphidae, members of the Australian genus Antipodogomphus generally posses a subcostal antenodal crossvein proximal to the first primary antenodal Ax1 in at least the fore wings and often in the hind wings (Theischinger & Hawking 2006).

<==Gomphoidea
    |--ProterogomphidaeRP02
    |--IctinogomphusWO91 [Ictinogomphinae, LindeniidaeTH06]
    |    |--I. australis (Selys 1873)TH06
    |    |--I. dobsoni (Watson 1969)TH06
    |    `--I. paulini Watson 1991TH06
    `--Gomphidae [Gomphinae, Gomphinidae]TH06
         |--Armagomphus armiger (Tillyard 1913)TH06
         |--Odontogomphus donnellyi Watson 1991TH06
         |--CacoidesBU02
         |--HageniusA07
         |--DromogomphusA07
         |--ErpetogomphusP74
         |--Gomphoides occultusP74, P92
         |--LanthusA07
         |--ProgomphusP74
         |--ArigomphusA07
         |--GomphurusA07
         |--HylogomphusA07
         |--StylurusA07
         |--Sinogomphus taushanensisRJ93
         |--OphiogomphusTW05
         |    |--O. serpentinusK01
         |    `--O. severusTW05
         |--ZephyrogomphusTH06
         |    |--Z. lateralis (Selys 1873)TH06
         |    `--Z. longipositor (Watson 1991)TH06
         |--IctinusSL89
         |    |--I. decoratusSL89
         |    `--I. melaenopsSL89
         |--GomphusFB03
         |    |--G. lucasii de Selys 1847E12
         |    |--G. spicatusA07
         |    `--G. vulgatissimusFB03
         |--MacrogomphusSL89
         |    |--M. albardaeSL89
         |    |--M. decemlineatusSL89
         |    `--M. quadratusSL89
         |--AustroepigomphusTH06
         |    |--A. (Austroepigomphus)TH06
         |    |    |--A. (A.) melaleucae (Tillyard 1909)TH06
         |    |    `--A. (A.) praeruptus (Selys 1857)TH06
         |    `--A. (Xerogomphus)TH06
         |         |--A. (X.) gordoni (Watson 1962)TH06
         |         `--A. (X.) turneri (Martin 1901)TH06
         |--AntipodogomphusTH06
         |    |--A. acolythus (Martin 1901)TH06
         |    |--A. dentosus Watson 1991TH06
         |    |--A. edentulus Watson 1991TH06
         |    |--A. hodgkini Watson 1969TH06
         |    |--A. neophytus Fraser 1958TH06
         |    `--A. proselythus (Martin 1901)TH06
         |--HemigomphusTH06
         |    |--H. atratus Watson 1991TH06
         |    |--H. comitatus (Tillyard 1909)TH06
         |    |--H. cooloola Watson 1991TH06
         |    |--H. gouldii (Selys 1854)TH06
         |    |--H. heteroclytus Selys 1854TH06
         |    |--H. magela Watson 1991TH06
         |    `--H. theischingeri Watson 1991TH06
         `--AustrogomphusTH06
              |  i. s.: A. lateralisP05
              |--A. (Austrogomphus)TH06
              |    |--A. (A.) angelorum Tillyard 1913TH06
              |    |--A. (A.) arbustorum Tillyard 1906TH06
              |    |--A. (A.) australis Dale 1854TH06
              |    |--A. (A.) collaris Hagen 1854TH06
              |    |--A. (A.) cornutus Watson 1991TH06
              |    |--A. (A.) doddi Tillyard 1909TH06
              |    |--A. (A.) guerini (Rambur 1842)TH06
              |    |--A. (A.) mjobergi Sjöstedt 1917TH06
              |    |--A. (A.) mouldsorum Theischinger 1999TH06
              |    |--A. (A.) ochraceus (Selys 1869)TH06
              |    `--A. (A.) pusillus Sjöstedt 1917TH06
              `--A. (Pleiogomphus)TH06
                   |--A. (P.) amphiclitus (Selys 1873)TH06
                   |--A. (P.) bifurcatus Tillyard 1909TH06
                   |--A. (P.) divaricatus Watson 1991TH06
                   `--A. (P.) prasinus Tillyard 1906TH06

*Type species of generic name indicated

References

[A07] Anderson, T. M. 2007. An assessment of water mite parasitism of dragonflies based on museum collections. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 61–69. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[BU02] Bechly, G., & K. Ueda. 2002. The first fossil record and first New World record for the dragonfly clade Chlorogomphida (Insecta: Odonata: Anisoptera: Araripechlorogomphidae n. fam.) from the Crato Limestone (Lower Cretaceous, Brazil). Stuttgarter Beiträge zur Naturkunde Serie B (Geologie und Paläontologie) 328: 1–11.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[FB03] Fleck, G., G. Bechly, X. Martínez-Delclòs, E. Jarzembowski, R. Coram & A. Nel. 2003. Phylogeny and classification of the Stenophlebioptera (Odonata: Epiproctophora). Annales de la Societe Entomologique de France 39 (1): 55–93.

[K01] Klapálek, F. 1901. Neuropteroidák [Neuropteroiden]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 203–221. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[P74] Paulson, D. R. 1974. Reproductive isolation in damselflies. Systematic Zoology 23 (1): 40–49.

[P05] Pinder, A. M. 2005. A review of biodiversity in wetlands with organic sediments on the Swan Coastal Plain, Western Australia, with an emphasis on aquatic invertebrates. Journal of the Royal Society of Western Australia 88 (3): 129–132.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

[RP02] Rasnitsyn, A. P., & L. N. Pritykina. 2002. Superorder Libellulidea Laicharting, 1781. Order Odonata Fabricius, 1792. The dragonflies. In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 97–104. Kluwer Academic Publishers: Dordrecht.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

[SL89] Selys Longchamps, E. de. 1889. Odonates de Sumatra comprenant les espèces recueillies à Pulo Nias par M. le D.r E. Modigliani. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 444–484.

[TW05] Terry, M. D., & M. F. Whiting. 2005. Mantophasmatodea and phylogeny of the lower neopterous insects. Cladistics 21: 240–257.

[TH06] Theischinger, G., & J. Hawking. 2006. The Complete Field Guide to Dragonflies of Australia. CSIRO Publishing: Collingwood (Victoria).

[WO91] Watson, J. A. L., & A. F. O’Farrell. 1991. Odonata (dragonflies and damselflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2nd ed. vol. 1 pp. 294–310. Melbourne University Press: Carlton (Victoria).

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