Gonyleptidae

Neopucroliella pertyi, copyright Luis Acosta.

Belongs within: Laminata.
Contains: Gonyassamiinae, Bourguyiinae, Tricommatinae, Mitobatinae, Hernandariinae, Caelopyginae, Progonyleptoidellinae, Sodreaninae, Discocyrtus, Eusarcus, Goniosomatinae, Gonyleptes, Geraecormobius, Metagonyleptes, Mischonyx, Heteropachylinae, Sadocus, Pachyloides.

The Gonyleptidae are a diverse group of harvestmen found in Central and South America, many of which are ornately armoured and/or brightly coloured. The family is divided between several subfamilies, though some of these (particularly the larger subfamilies Gonyleptinae and Pachylinae) remain poorly defined.

Sphaerobunus is a genus of gonyleptid harvestmen found in southern Brazil, males of which possess a large transverse apical apophysis on coxa IV and a comb of very long spines on femur IV.

Characters (from Kury & Pinto-da-Rocha 2007): Carapace usually much narrower than opisthosomal scutum, body rarely subrectangular. Frontal hump usually present; with tubercles short or long. Dorsal scutum usually longer than wide. With one or two ozopores on each side. Body depressed or convex. Common ocularium always present. Three or four areas on dorsal scutum; sulci well marked; area I divided by median groowve; IV rarely divided. Chelicerae usually similar in size and small in both sexes, rarely hypertelic in male. Coxa IV usually dorsally visible in entire extension; with dorsoapical apophysis short to robust, single or bifid, much more developed on males. Trochanter, femur and tibia IV often armed with tubercles/apophyses in males. Scopulae usually present on legs III and IV. Claws usually smooth, sometimes pectinate. Penis with ventral plate well defined, rectangular or pyriform with basal lobes more or less developed; lateral margin of ventral plate with two groups of setae; glans columnar, more or less elongate, rarely with dorsal process, commonly with ventral process.

Gonyleptidae [Ejonyleptidae, Golyleptidae, Gonileptidae, Gonyleptoidae]
    |--Ampycinae [Ampycidae]A19
    |    |  i. s.: Hexabunus Roewer 1913K03b
    |    |           `--*H. armillatus (Butler 1873) [=Gonyleptes armillatus]K03b
    |    |         Ampycella Roewer 1929KA-Z11
    |    |           `--*A. spiniventris Roewer 1929 [=Nesopachylus spiniventris]K03b
    |    |         Glysterus Roewer 1931KA-Z11 (see below for synonymy)
    |    |           |--*G. scutatus Roewer 1931 [incl. *Rooma caudaspina Goodnight & Goodnight 1942]K03b
    |    |           |--G. calcartibialis (Roewer 1943) [=*Triaenoplus calcartibialis]K03b
    |    |           |--G. costaricensis Roewer 1943 [=*Glysteroides costaricensis]K03b
    |    |           |--G. guatemalensis (Roewer 1943) [=Glysteroides guatemalensis]K03b
    |    |           |--G. laeviscutatus (Roewer 1943) [=*Phalacrobunus laeviscutatus]K03b
    |    |           `--G. metatarsalis (Roewer 1943) [=*Angathisoma metatarsalis]K03b
    |    |         Parahernandria Goodnight & Goodnight 1947KA-Z11
    |    |           |--*P. ventralis (Banks 1914)K03b, KA-Z11 [=Hernandria (l. c. for Hernandaria) ventralisK03b]
    |    |           `--P. spinosa (Banks 1909)KA-Z11 [=Hernandria (l. c. for Hernandaria) spinosaK03b]
    |    |         Neopachyloides Roewer 1913KA-Z11
    |    |           `--*N. spinipes Roewer 1913K03b
    |    |         Nesopachylus Chamberlin 1925KA-Z11 (see below for synonymy)
    |    |           |--*N. monoceros Chamberlin 1925 [=N. monoceras (l. c.)]K03b
    |    |           `--N. maculatus (Roewer 1932) [=*Gorgonasta maculata]K03b
    |    |         Sibollus Roewer 1929KA-Z11
    |    |           `--*S. margaritatus Roewer 1929K03b [=Neopachyloides margaritatusKA-Z11]
    |    |         Thaumatopachylus Roewer 1929KA-Z11
    |    |           `--*T. setulosus Roewer 1929K03b [=Neopachyloides setulosusKA-Z11]
    |    |--Ampycus Simon 1879A19, K03b [incl. Itequahy Mello-Leitão 1949K03b]
    |    |    |--*A. telifer (Butler 1873) [=Gonyleptes telifer; incl. *Itequahy ensiferus Mello-Leitão 1949]K03b
    |    |    `--A. granulosus (Soares 1970) [=Bullaepus granulosus]K03b
    |    `--+--Hutamaia Soares & Soares 1977A19, KA-Z11
    |       |    `--*H. caramaschii Soares & Soares 1977K03b
    |       `--Licornus Roewer 1932A19, K03b
    |            |--*L. perfectus Roewer 1932K03b
    |            |--L. atroluteus Roewer 1959K03b
    |            `--L. tama Villareal & Kury 2012A19
    `--+--TricommatinaeA19
       `--+--+--MitobatinaeDG09
          |  `--+--+--+--+--HernandariinaeDG09
          |     |  |  |  `--CaelopyginaeDG09
          |     |  |  `--+--ProgonyleptoidellinaeDG09
          |     |  |     `--SodreaninaeDG09
          |     |  `--+--+--DiscocyrtusM11
          |     |     |  `--+--EusarcusA19
          |     |     |     `--Eubalta Roewer 1923A19, K03b [=Balta Sørensen 1902 non Tapper 1893K03b, Baltanata Strand 1942K03b]
          |     |     |          `--E. planiceps (Gervais 1842)KM20 (see below for synonymy)
          |     |     `--+--GoniosomatinaeDG09
          |     |        `--Mangaratiba Mello-Leitão 1940M11, K03 [incl. Canopilio Soares 1968K03]
          |     |             |--*M. monstrosa Mello-Leitão 1940K03
          |     |             |--M. acanthoproctus (Soares 1968)KA-Z11 (see below for synonymy)
          |     |             `--M. angulispinosis (Soares 1966)KA-Z11 [=Piresa angulispinosisK03, M. angulispinosaK03]
          |     `--GonyleptinaeDG09
          |          |--GonyleptesP-R02
          |          |--GeraecormobiusK03b
          |          |--MetagonyleptesK03b
          |          |--MischonyxK03b
          |          |--Haversia Roewer 1913K03b
          |          |    `--*H. defensa (Butler 1876) [=Gonyleptes defensus]K03b
          |          |--Progonyleptes Roewer 1913K03b
          |          |    `--*P. borellii (Sørensen 1895) [=Gonyleptes borellii]K03b
          |          |--Eugonyleptes Roewer 1913K03b
          |          |    `--*E. scaber (Kirby 1819)K03b, KA-Z11 [=Gonyleptes scaberK03b, Discocyrtus scaberK03b]
          |          |--Triaenomeros Roewer 1913K03b
          |          |    `--*T. olivaceum Roewer 1913 [=T. olivaceus]KM20
          |          |--Adhynastes Roewer 1930K03b
          |          |    `--*A. tenuis Roewer 1930K03b
          |          |--Cearinus Roewer 1929 [incl. Nemoribalta Mello-Leitão 1941]K03b
          |          |    `--*C. corniger Roewer 1929 [incl. *Nemoribalta plana Mello-Leitão 1941]K03b
          |          |--Centroleptes Roewer 1943K03b
          |          |    `--*C. flavus Roewer 1943K03b
          |          |--Deltaspidium Roewer 1927K03b
          |          |    `--D. asper (Perty 1833) (see below for synonymy)K03b
          |          |--Gonazula Roewer 1930K03b
          |          |    `--*G. gibbosa Roewer 1930K03b
          |          |--Guatubesia Soares 1978K03b
          |          |    `--*G. clarae Soares 1978K03b
          |          |--Gyndesops Roewer 1943K03b
          |          |    `--*G. denisi Roewer 1943 [=Temucus denisi]K03b
          |          |--Hoggellula Roewer 1930K03b
          |          |    `--*H. vallentini (Hogg 1913) [=Sadocus vallentini]K03b
          |          |--Holoversia Mello-Leitão 1940K03b
          |          |    `--*H. nigra Mello-Leitão 1940K03b
          |          |--Proctobunoides Mello-Leitão 1944K03b
          |          |    `--*P. tuberosus Mello-Leitão 1944K03b
          |          |--Stefanesia Soares & Soares 1988K03b
          |          |    `--*S. avramae Soares & Soares 1988K03b
          |          |--Urodiabunus Mello-Leitão 1935K03b
          |          |    `--*U. arlei Mello-Leitão 1935K03b
          |          |--Ypsilonurus Mello-Leitão 1933K03b
          |          |    `--*Y. mutilatus Mello-Leitão 1933K03b
          |          |--Progonyleptoides Roewer 1917 [incl. Paranabius Roewer 1943]K03b
          |          |    |--*P. spinifrons Roewer 1917K03b
          |          |    `--P. castaneus (Roewer 1943) [=*Paranabius castaneus]K03b
          |          |--Currala Roewer 1927K03b
          |          |    |--*C. spinifrons Roewer 1927K03b
          |          |    `--C. bahiensis Soares 1972K03b
          |          |--Schenkelibunus Strand 1932K03b, KA-Z11 (see below for synonymy)
          |          |    |--*S. perditus (Mello-Leitão 1927)K03b, KA-Z11 [=*Hanseniella perditaK03b, Friburgoia perditaK03b]
          |          |    `--S. impar (Mello-Leitão 1932)KA-Z11 (see below for synonymy)
          |          |--Gonyleptellus Roewer 1930 [incl. Stephanocranion Mello-Leitão 1931]K03b
          |          |    |--G. cancellatus (Roewer 1917) (see below for synonymy)K03b
          |          |    `--G. bufoninus (Mello-Leitão 1949) [=Stephanocranion bufoninus]K03b
          |          |--Inhuma Piza 1938 [incl. Hugoesia Soares 1968]K03b
          |          |    |--*I. pessoai Piza 1938K03b
          |          |    `--I. lopesi (Soares 1968) [=*Hugoesia lopesi]K03b
          |          |--Uracantholeptes Mello-Leitão 1926 [incl. Gonypernoides Mello-Leitão 1932]K03b
          |          |    |--*U. anomalus (Mello-Leitão 1922) (see below for synonymy)K03b
          |          |    `--U. damicoi (Soares & Soares 1945) [=Gonypernoides damicoi]K03b
          |          |--Megapachylus Roewer 1913 (see below for synonymy)K03b
          |          |    |--*M. grandis Roewer 1913 (see below for synonymy)K03b
          |          |    |--M. mutilatus (Mello-Leitão 1932)K03b [=Gonyperna mutilataK03b, Paragonyleptes mutilatusSS49]
          |          |    `--M. simoni (Roewer 1930) (see below for synonymy)K03b
          |          |--Tupacarana Mello-Leitão 1939K03b
          |          |    |--*T. serrina Mello-Leitão 1939K03b
          |          |    |--T. gofferjei Soares & Soares 1947K03b
          |          |    `--T. marmorata Mello-Leitão 1939K03b
          |          |--Juticus Roewer 1943KA20
          |          |    |--*J. furcidens Roewer 1943K03b
          |          |    |--J. calcaratus (Roewer 1930) [=Paragonyleptes calcaratus]KA20
          |          |    |--J. frontalis (Roewer 1930) [=Moreira frontalis, Paragonyleptes frontalis]KA20
          |          |    `--J. hamiferus (Mello-Leitão 1935)KA20 [=Gonyleptilus hamiferusKA20, Paragonyleptes hamiferusK03b]
          |          |--Neosadocus Mello-Leitão 1926K-PR07, K03b (see below for synonymy)
          |          |    |--*N. bufo (Mello-Leitão 1923) (see below for synonymy)K03b
          |          |    |--N. maximus (Giltay 1928) [=Neomitobates maximus, *Mitoperna maxima]K03b
          |          |    |--N. misandrus (Mello-Leitão 1934) [=Metagonyleptes misandrus]K03b
          |          |    |--N. robustus (Mello-Leitão 1936) [=*Ilhania robusta]K03b
          |          |    `--N. variabilis (Mello-Leitão 1935)DG09 [=*Bunoweyhia variabilisK03b]
          |          |--Liogonyleptoides Mello-Leitão 1925 (see below for synonymy)K03b
          |          |    |--*L. inermis (Mello-Leitão 1922) (see below for synonymy)K03b
          |          |    |--L. capichaba Soares & Soares 1946K03b
          |          |    |--L. heliae Kury 2003 [=L. minensis Soares 1966 non Chaquesia minensis Piza 1946]K03b
          |          |    |--L. minensis (Piza 1946) [=Chaquesia minensis]K03b
          |          |    `--L. tetracanthus (Mello-Leitão 1932) [=*Pachyleptes tetracanthus]K03b
          |          |--Parampheres Roewer 1913KP-R07, K03b (see below for synonymy)
          |          |    |--*P. pectinatus Roewer 1913K03b
          |          |    |--P. bimaculatus (Mello-Leitão 1932) (see below for synonymy)K03b
          |          |    |--P. boliviensis (Roewer 1931) [=*Callampheres boliviensis]K03b
          |          |    |--P. lucidus (Mello-Leitão 1940) [=Penygorna lucida, Arleius lucidus, Ilhaia lucida]K03b
          |          |    |--P. ronae (Mello-Leitão 1927) (see below for synonymy)K03b
          |          |    `--P. tibialis Roewer 1917 [incl. Philocrates maculatus Sørensen in Roewer 1943 (n. n.)]K03b
          |          `--Sphaerobunus Roewer 1917DG09, K03b (see below for synonymy)
          |               |--*S. rhinoceros Roewer 1917K03b
          |               |--S. editus (Roewer 1943) (see below for synonymy)K03b
          |               |--S. fallax (Mello-Leitão 1932) (see below for synonymy)K03b
          |               |--S. fulvigranulatus (Mello-Leitão 1922) (see below for synonymy)K03b
          |               |--S. marmoratus (Mello-Leitão 1940) [=Progonyleptes marmoratus]K03b
          |               |--S. pictus (Piza 1942) (see below for synonymy)K03b
          |               |--S. singularis (Mello-Leitão 1935) (see below for synonymy)K03b
          |               `--S. variolosus (Mello-Leitão 1940) (see below for synonymy)K03b
          `--+--+--HeteropachylinaeM11
             |  `--Cobania Roewer 1913M11, K03b (see below for synonymy)
             |       |--*C. picea (Bertkau 1880) (see below for synonymy)K03b
             |       `--C. validissima (Mello-Leitão 1933) [=*Flangeia validissima]K03b
             `--PachylinaeDG09
                  |--SadocusP-R02
                  |--PachyloidesK08
                  |--Parapucrolia Roewer 1917 [incl. Goytacazia Mello-Leitão 1942, Goitacazia (l. c.), Goytacazio (l. c.)]K03b
                  |    `--*P. ocellata Roewer 1917 (see below for synonymy)K03b
                  |--Meteusarcus Roewer 1913K03b
                  |    `--*M. armatus Roewer 1913K03b
                  |--Paraphalangodus Roewer 1915K03b
                  |    `--*P. synacanthus Roewer 1915K03b
                  |--Acanthopachylus Roewer 1913 (see below for synonymy)K03b
                  |    `--*A. aculeatus (Kirby 1818)K03b, KA-Z11 (see below for synonymy)
                  |--Allogonyleptes Roewer 1917K03b
                  |    `--*A. insignitus Roewer 1917K03b
                  |--Anoplogynopsis Soares 1966K03b
                  |    `--*A. concolor Soares 1966K03b
                  |--Antetriceras Roewer 1949K03a
                  |    `--*A. signatum Roewer 1949K03b, KA-Z11, K03b [=A. signatusK03b]
                  |--Berlaia Mello-Leitão 1940K03b
                  |    `--*B. dissimilis Mello-Leitão 1940 (see below for synonymy)K03b
                  |--Biconisoma Roewer 1936K03b
                  |    `--*B. mirabile Roewer 1936K03b, KA-Z11, K03b [=B. mirabilisK03b]
                  |--Bullaepus Roewer 1930 [=Bullaeepus (l. c.)]K03b
                  |    `--*B. enoplus (Chamberlin 1916) [=Gonoleptes (l. c. for Gonyleptes) enoplus]K03b
                  |--Bunoplus Roewer 1927K03b
                  |    `--*B. pachypalpis Roewer 1927 [=B. pachypalpi (l. c.)]K03b
                  |--Calcarogyndes Mello-Leitão 1932 [=Calcogyndes (l. c.), Callcarogyndes (l. c.)]K03b
                  |    `--*C. calcar (Roewer 1913) [=Metagyndes calcar]K03b
                  |--Caldanatus Roewer 1943K03b
                  |    `--*C. marginalis Roewer 1943K03b
                  |--Camposicoloides Soares 1944K03b
                  |    `--*C. mendax Soares 1944K03b
                  |--Capichabesia Soares 1944K03b
                  |    `--*C. rarissima Soares 1944K03b
                  |--Ceratoleptes Soares & Soares 1979K03b
                  |    `--*C. proboscis Soares & Soares 1979 (see below for synonymy)K03b
                  |--Chaquesia Soares 1944K03b
                  |    `--*C. guaranitica (Mello-Leitão 1933) [=Goyazella guaranitica]K03b
                  |--Chauveaua Canals 1939K03b
                  |    `--*C. nahuelhapiensis Canals 1939 [=Phalangodus nahuelhapiensis]K03b
                  |--Chilebalta Roewer 1961K03b
                  |    `--*C. angulipes Roewer 1961K03b
                  |--Chilegyndes Roewer 1961K03b
                  |    `--*C. phillipsoni Roewer 1961K03b
                  |--Corralia Roewer 1913K03b
                  |    `--*C. depressa (Loman 1899) [=Gonyleptes depressus; incl. Neogonyleptes ignotus Soares 1968]K03b
                  |--Ergastria Mello-Leitão 1941 [=Castellanosia Mello-Leitão 1937 non Kraglievitch 1932]K03b
                  |    `--*E. forficula (Mello-Leitão 1937) [=*Castellanosia forficula]K03b
                  |--Fonckia Roewer 1913K03b
                  |    `--*F. processigera (Sørensen 1902) [=Gonyleptes processiger, F. prossigera (l. c.)]K03b
                  |--Giupponia Pérez G. & Kury 2002K03b
                  |    `--*G. chagasi Pérez-González & Kury 2002K03b
                  |--Goodnightiella Soares & Soares 1945 [incl. Prosampycoides Soares & Bauab-Vianna 1972]K03b
                  |    `--*G. impar Soares & Soares 1945 [incl. *Prosampycoides spinipes Soares & Bauab-Vianna 1972]K03b
                  |--Gyndesoides Mello-Leitão 1933K08 [incl. Klobinia Mello-Leitão 1936K03b]
                  |    `--*G. dispar Mello-Leitão 1933 [incl. *Klobinia infuscata Mello-Leitão 1936]K03b
                  |--Harpachylus Roewer 1943K03b
                  |    `--*H. tibialis Roewer 1943K03b
                  |--Huadquina Roewer 1930K03b
                  |    `--*H. huadquinae (Chamberlin 1916) (see below for synonymy)K03b
                  |--Hyperpachylus Roewer 1957K03b
                  |    `--*H. mirabilis Roewer 1957K03b
                  |--Ibarra Roewer 1925K03b
                  |    `--*I. festae Roewer 1925K03b
                  |--Iguassua Mello-Leitão 1935K03b
                  |    `--*I. humilis Mello-Leitão 1935K03b
                  |--Iguassoides Soares & Soares 1954K03b
                  |    `--*I. lucidus (Soares 1944) [=Iguassua lucida]K03b
                  |--Itatiaincola Soares & Soares 1948K03b
                  |    `--*I. nanus Soares & Soares 1948K03b
                  |--Izecksohnopilio Soares 1977K03b
                  |    `--*I. eugenioi Soares 1977K03b
                  |--Junicus Goodnight & Goodnight 1947 [incl. Sokkupia Mello-Leitão 1949]K03b
                  |    `--*J. gerhardi Goodnight & Goodnight 1947 (see below for synonymy)K03b
                  |--Maromba Soares & Soares 1954K03b
                  |    `--*M. dandrettai Soares & Soares 1954K03b
                  |--Metadiscocyrtus Roewer 1929K03b
                  |    `--M. fornicatus (Sørensen 1884) [=Discocyrtus fornicatus]K03b
                  |--Metagyndoides Mello-Leitão 1931K03b
                  |    `--*M. granulatus Mello-Leitão 1931 [=Metagyndes granulata, Metagyndes granulatus]K03b
                  |--Metaphalangodella Roewer 1915K03b
                  |    `--*M. pachyliformis Roewer 1915K03b
                  |--Oliverius Soares & Soares 1945K03b
                  |    `--*O. jordanensis Soares & Soares 1945K03b
                  |--Pachylusius Mello-Leitão 1949K08
                  |    `--*P. incus Mello-Leitão 1949K03b
                  |--Paraprosontes Soares & Soares 1947K03b
                  |    `--*P. inermis Soares & Soares 1947K03b
                  |--Pareusarcus Roewer 1929K03b
                  |    `--*P. corniculatus Roewer 1929K03b
                  |--Passosa Roewer 1928K03b, KA-Z11
                  |    `--*P. instructa Roewer 1928K03b, KA-Z11
                  |--Pichitus Roewer 1959K03b
                  |    `--*P. aniarmatus Roewer 1959K03b
                  |--Pirunipygus Roewer 1936K03b
                  |    `--*P. paradoxus Roewer 1936K03b
                  |--Planiphalangodus Roewer 1929K03b
                  |    `--*P. robustus Roewer 1929 [=Pachyloides robustus]K03b
                  |--Polyacanthoprocta Mello-Leitão 1927K03b
                  |    `--*P. orina (Chamberlin 1916) [=Pachylus orinus]K03b
                  |--Pseudoacrographinotus Soares 1966K03b
                  |    `--*P. simplex Soares 1966K03b
                  |--Punagraphinotus Soares & Bauab-Vianna 1973K03b, KA-Z11
                  |    `--*P. punae Soares & Bauab-Vianna 1973K03b, KA-Z11
                  |--Riosegundo Canals 1943K03b
                  |    `--*R. birabeni Canals 1943K03b
                  |--Schubartesia Soares 1944K03b
                  |    `--*S. singularis Soares 1944K03b
                  |--Tarmapachylus Roewer 1956K03b [=Tarmopachylus (l. c.)KA-Z11]
                  |    `--*T. koepckei Roewer 1956K03b
                  |--Temucus Roewer 1943K03b
                  |    `--*T. palpiconus Roewer 1943K03b
                  |--Tingomaria Mello-Leitão in Mello-Leitão & Feio 1949 1949K03b, KM20 (see below for synonymy)
                  |    `--*T. hydrophila Mello-Leitão in Mello-Leitão & Feio 1949KM20 (see below for synonymy)
                  |--Unduavius Roewer 1929K03b
                  |    `--*U. ornatus Roewer 1929K03b
                  |--Victoriaincola Soares & Soares 1946K03b
                  |    `--*V. penhae Soares & Soares 1946K03b
                  |--Spinivunus Roewer 1943K03b
                  |    `--*S. adumbratus Roewer 1943K03b
                  |--Bristoweia Mello-Leitão 1924K08 [=Bristoweria (l. c.)K03b]
                  |    `--*B. diamantinae Mello-Leitão 1924K03b
                  |--Eopachylus Mello-Leitão 1931 [=Eupachylus (l. c.)]K03b
                  |    `--*E. ignotus Mello-Leitão 1931K03b
                  |--Paraluederwaldtia Mello-Leitão 1927K03b
                  |    `--*P. bituberculata (Mello-Leitão 1922) (see below for synonymy)K03b
                  |--Pherania Strand 1942 [=Phera Sørensen 1932 non Stål 1864, Phaera (l. c.)]K03a
                  |    |--*P. pygmaea (Sørensen 1932) [=*Phera pygmaea]K03a
                  |    `--P. giupponii Kury 2003K03a
                  |--Soaresia Soares 1945K08
                  |    |--*S. uncina Soares 1945K08
                  |    `--S. forficula Kury 2008K08
                  |--Acanthoprocta Loman 1899 (see below for synonymy)K03b
                  |    |--*A. pustulata Loman 1899 (see below for synonymy)K03b
                  |    `--A. conica Maury 1991K03b
                  |--Parabalta Roewer 1913KP-R07, K03b
                  |    |--*P. reedii (Butler 1876) [=Gonyleptes reedii, Lycomedes reedii]K03b
                  |    `--P. cristobalia (Roewer 1943) [=Neopucroliella cristobalia]K03b
                  |--Metapachyloides Roewer 1917K03b
                  |    |--*M. rugosus Roewer 1917K03b
                  |    `--M. almeidai Soares & Soares 1946K03b
                  |--Camposicola Mello-Leitão 1924K08
                  |    |--*C. altifrons Mello-Leitão 1924K03b
                  |    `--C. sanctateresae (Soares & Soares 1946) [=Camposicoloides sanctateresae]K03b
                  |--Carlotta Roewer 1943 (see below for synonymy)K03b
                  |    |--*C. serratipes Roewer 1943 (see below for synonymy)K03b
                  |    `--C. dubia (Soares 1944) [=*Pseudoneogonyleptoides dubius]K03b
                  |--Ceropachylinus Mello-Leitão 1943K03b
                  |    |--*C. granulosus Mello-Leitão 1943K03b
                  |    `--C. peruvianus Roewer 1956K08
                  |--Diconospelta Canals 1934K03b
                  |    |--*D. gallardoi Canals 1934K03b
                  |    `--D. vazferreirae Mello-Leitão 1946K03b
                  |--Discocyrtulus Roewer 1927K03b
                  |    |--*D. bresslaui Roewer 1927K03b
                  |    `--D. marginalis Roewer 1929K03b
                  |--Gyndoides Mello-Leitão 1927K03b
                  |    |--*G. elaphus Mello-Leitão 1927K03b
                  |    `--G. springmanni Soares & Soares 1947K03b
                  |--Huralvioides Soares 1970K03b
                  |    |--*H. paruensis Soares 1970K03b
                  |    `--H. hoeferi Kury 1995K03b
                  |--Metalycomedes Mello-Leitão 1927K03b
                  |    |--*M. perditus Mello-Leitão 1927K03b
                  |    `--M. secundus Soares & Soares 1947K03b
                  |--Meteusarcoides Mello-Leitão 1922 [incl. Cercopachylus Piza 1940]K03b
                  |    |--*M. mutilatus Mello-Leitão 1922 [incl. Uropachylus grovesi Soares 1944]K03b
                  |    `--M. caudatus (Piza 1940) (see below for synonymy)K03b
                  |--Palcapachylus Roewer 1952K03b
                  |    |--*P. bicalcariventris Roewer 1952K03b
                  |    `--P. peruvianus Roewer 1959K03b
                  |--Kuryella Özdikmen 2006O06 [=Paranaleptes Soares & Soares 1947 non Breuning 1937K03b, O06]
                  |    |--*K. xanthoacantha (Soares & Soares 1947) [=*Paranaleptes xanthoacanthus]O06
                  |    `--K. melanoacantha (Soares & Soares 1947)O06 (see below for synonymy)
                  |--Pseudogyndes Mello-Leitão 1932K03b
                  |    |--*P. subsimilis (Roewer 1913) [=Metagyndes subsimilis]K03b
                  |    `--P. marginatus Roewer 1961K03b, KA-Z11 [=P. marginataK03b]
                  |--Tumbesia Loman 1899K03b
                  |    |--*T. fuliginosa Loman 1899K03b
                  |    `--T. aculeata Roewer 1930K03b
                  |--Yraguara Mello-Leitão 1937 [incl. Melloleitanius Piza 1938]K03b
                  |    |--*Y. annulipes Mello-Leitão 1937 [incl. *Melloleitanius barretensis Piza 1938, Y. barretensis]K03b
                  |    `--Y. fleuryi Soares 1943K03b
                  |--Iandumoema Pinto-da-Rocha 1997K03b, KA-Z11
                  |    |--*I. uai Pinto-da-Rocha 1996K03b, A19
                  |    |--I. setimapocu Hara & Pinto-da-Rocha 2008P-GC17
                  |    `--I. smeagol Pinto-da-Rocha, Fonseca-Ferreira & Bichuette 2015P-GC17
                  |--Ceropachylus Mello-Leitão 1942 [incl. Allelochirus Mello-Leitão 1942, Pachylla Roewer 1959]K03b
                  |    |--*C. frizzellae Mello-Leitão 1942K03b
                  |    |--C. areolatus (Roewer 1959) [=*Pachylla areolata]K03b
                  |    `--C. singularis (Mello-Leitão 1942) [=*Allelochirus singularis]K03b
                  |--Huasampillia Roewer 1913 [=Huasampilla (l. c.); incl. Lucma Roewer 1930]K03b
                  |    |--*H. terribilis (Butler 1876) [=Gonyleptes terribilis]K03b
                  |    |--H. albipustulata Roewer 1943K03b
                  |    `--H. scotia (Chamberlin 1916) [=Gonyleptes scotius, *Lucma scotia]K03b
                  |--Hypophyllonomus Giltay 1928 [incl. Parabristoweia Piza 1938]K03b
                  |    |--*H. longipes Giltay 1928K03b
                  |    |--H. callidus Soares 1966K03b
                  |    `--H. maculipalpi (Piza 1938) [=*Parabristoweia maculipalpi]K03b
                  |--Ogloblinia Canals 1933 (see below for synonymy)K03b
                  |    |--*O. loretoensis Canals 1933 (see below for synonymy)K03b
                  |    |--O. argenteopilosa (Mello-Leitão 1935) (see below for synonymy)K03b
                  |    `--O. pulchra Pinto-da-Rocha & Soares 1995K03b
                  |--Paradiscocyrtus Mello-Leitão 1927K03b
                  |    |--*P. neglectus Mello-Leitão 1927K03b
                  |    |--P. cerayanus Roewer 1929K03b
                  |    `--P. trochanteralis Roewer 1929K03b
                  |--Pseudogyndesoides Soares 1944K03b
                  |    |--*P. latus Soares 1944K03b
                  |    |--P. bariguiensis Soares 1945K03b
                  |    `--P. pallidus Soares & Soares 1945K03b
                  |--Singram Mello-Leitão 1937K03b
                  |    |--*S. simplex Mello-Leitão 1937K03b
                  |    |--S. brasiliensis (Mello-Leitão 1940) [=Oglobinia brasiliensis]K03b
                  |    `--S. singularis Soares & Soares 1948K03b
                  |--Ubatubesia Soares 1945 [incl. Ubatubesiopsis Soares & Soares 1954]K03b
                  |    |--*U. oliverioi Soares 1945 [incl. U. travassoi Soares & Soares 1947]K03b
                  |    |--U. amplicoxae Pinto-da-Rocha 2000K03b
                  |    `--U. rabelloi (Soares & Soares 1954) [=*Ubatubesiopsis rabelloi]K03b
                  |--Punrunata Roewer 1952K03b
                  |    |--*P. tibialis Roewer 1952K03b
                  |    |--P. brasiliensis Soares & Soares 1979K03b
                  |    `--P. femoralis Soares & Bauab-Vianna 1972K03b
                  |--Lacronia Strand 1942K08 [=Luederwaldtia Mello-Leitão 1923 non Schmidt 1922KO06]
                  |    |--*L. serripes (Mello-Leitão 1923) [=*Luederwaldtia serripes]KO06
                  |    |--L. camboriu Kury 2003KO06
                  |    |--L. ceci Kury & Orrico 2006KO06
                  |    `--L. ricardoi Kury 2003KO06
                  |--Pachyloidellus Müller 1917K03b, KA-Z11 (see below for synonymy)
                  |    |--P. butleri (Thorell 1877) (see below for synonymy)K03b
                  |    |--P. borelliiCM07
                  |    |--P. fulvigranulatus (Mello-Leitão 1930) (see below for synonymy)K03b
                  |    `--P. goliath Acosta 1993K03b
                  |--Pygophalangodus Mello-Leitão 1931K03b (see below for synonymy)
                  |    |--*P. gemignanii Mello-Leitão 1931 [=P. geminganii (l. c.)]K03b
                  |    |    |--P. g. gemignaniiK03b
                  |    |    `--P. g. uruguayensis Ringuelet 1955K03b
                  |    `--P. canalsi (Mello-Leitão 1931) (see below for synonymy)K03b
                  |--Parapachyloides Roewer 1913 (see below for synonymy)K03b
                  |    |--*P. uncinatus (Sørensen 1879) [=Gonyleptes uncinatus, Pachyloides uncinatus]K03b
                  |    |--P. armatus (Mello-Leitão 1931) (see below for synonymy)K03b
                  |    |--P. bispinifrons (Roewer 1943) [=Liogyndulus bispinifrons]KP-R20
                  |    |--P. dentipes Roewer 1913K03b
                  |    `--P. fontanensis (Canals 1935) [=*Apophysigerus fontanensis, Goyazella fontanensis]K03b
                  |--Pachylibunus Roewer 1913K03b
                  |    |--*P. grandis Roewer 1913K03b
                  |    |--P. armatissimus Roewer 1917K03b
                  |    |--P. gomesianus Mello-Leitão 1922K03b
                  |    |--P. hamatus Roewer 1930K03b
                  |    `--P. nigripes (Mello-Leitão 1942) [=Caldasius nigripes]KM20
                  |--Pucrolia Sørensen 1895K03b
                  |    |--*P. minuta (Sørensen 1884) [=Pachylus minutus]K03b
                  |    |--P. dubitata Soares & Soares 1945K03b
                  |    |--P. gracilipes (Canestrini 1888) (n. d.) [=Pachylus gracilipes]K03b
                  |    |--P. grandis Mello-Leitão 1930K03b
                  |    `--P. pulcherrima Soares 1966K03b
                  |--Metabalta Roewer 1913K03b
                  |    |--*M. tuberculata Roewer 1913K03b
                  |    |--M. albipes Mello-Leitão 1931 [incl. M. porteri Mello-Leitão 1936]K03b
                  |    |--M. efformata Roewer 1929K03b
                  |    |--M. geniculata Roewer 1929K03b
                  |    `--M. hostilis Roewer 1913K03b
                  |--Guaraniticus Mello-Leitão 1933 [incl. Guaranilia Mello-Leitão 1937]K03b
                  |    |--*G. lesserti Mello-Leitão 1933K03b
                  |    |--G. flavimaculatus Soares & Soares 1947K03b
                  |    |--G. nigrosulcatus (Mello-Leitão 1937) [=*Guaranilia nigrosulcata]K03b
                  |    |--G. olivaceus (Roewer 1913) [=Cobania olivacea]K03b
                  |    `--G. tetracalcar Soares & Soares 1945K03b
                  |--Neogonyleptes Roewer 1913 [incl. Neogonyleptoides Roewer 1913]K03b
                  |    |--*N. docilis (Butler 1876) [=Gonyleptes docilis]K03b
                  |    |--N. chilensis (Roewer 1913) [=Neogonyleptoides chilensis]K03b
                  |    |--N. frontalis (Sørensen 1902) [=Gonyleptes frontalis]K03b
                  |    |--N. hamatus Soares 1968K03b
                  |    `--N. karschii (Sørensen 1902) [=Gonyleptes karschii, *Neogonyleptoides karschii]K03b
                  |--Pachylus Koch 1839 [=Kochipachylus Strand 1942; incl. Acanthopachyloides Piza 1942]K03b
                  |    |--P. chilensis (Gray 1833)KM20 (see below for synonymy)
                  |    |--P. acanthops (Gervais 1849) [=Gonyleptes acanthops, G. acantops (l. c.)]K03b
                  |    |--P. crassus (Roewer 1943) [=Pachyloidellus crassus, Progyndes crassus]K03b
                  |    |--P. paessleri Roewer 1913K03b
                  |    |--P. quinamavidensis Muñoz-Cuevas 1969K03b
                  |    `--P. vachoni Muñoz-Cuevas 1970K03b
                  |--Metagraphinotus Mello-Leitão 1927 [incl. Jacarepaguana Mello-Leitão 1932]K03b
                  |    |--*M. catharinensis Mello-Leitão 1927K03b
                  |    |--M. arlei Mello-Leitão 1935K03b
                  |    |--M. berlae (Mello-Leitão 1932) [=*Jacarepaguana berlae]K03b
                  |    |--M. pectinifemur (Piza 1943) [=Jacarepaguana pectinifemur]K03b
                  |    |--M. sooretamae Soares & Soares 1946K03b
                  |    `--M. trochanterspinosus Soares & Soares 1947K03b
                  |--Uropachylus Mello-Leitão 1922 (see below for synonymy)K03b
                  |    |--*U. striatus Mello-Leitão 1922K03b
                  |    |--U. anthophilus (Mello-Leitão 1926) [=Ypiranga anthophila]K03b
                  |    |--U. gratiosus Soares & Soares 1954K03b
                  |    |--U. itatiaia Soares & Soares 1970K03b
                  |    |--U. nasutus Soares & Soares 1947 [=U. nasutis (l. c.)]K03b
                  |    `--U. ypiranga (Mello-Leitão 1922) (see below for synonymy)K03b
                  |--Acrographinotus Holmgren 1916A02, KA-Z11 (see below for synonymy)
                  |    |--*A. erectispina Roewer 1929A02
                  |    |--A. ceratopygus (Soares & Bauab-Vianna 1972) [=*Ctatoproceros ceratopygus]K03b
                  |    |--A. curvispina Roewer 1929A02
                  |    |--A. mitmaj Acosta 2002A02
                  |    |--A. niawpaq Acosta 2001A02
                  |    |--A. opacus (Roewer 1952) [=Liogyndulus opacus]KP-R20
                  |    `--A. ortizi (Roewer 1957)A02 [=*Liographinotus ortiziK03b, Sokkupia ortiziK03b]
                  |--Triglochinura Mello-Leitão 1924 (see below for synonymy)K03b
                  |    |--*T. curvispina Mello-Leitão 1924K03b (see below for synonymy)
                  |    |--T. ancilla (Mello-Leitão 1937) [=*Tayoca ancilla, Piresa ancilla]K03b
                  |    |--T. apiaiensis (Soares & Bauab-Vianna 1972) [=Piresa apiaiensis]K03b
                  |    |--T. indicta (Soares & Soares 1970) [=Piresa indicta]K03b
                  |    |--T. langei (Soares & Soares 1949) [=Piresa langei]K03b
                  |    |--T. timida (Mello-Leitão 1940) [=Itaoca timida]K03b
                  |    `--T. villosa (Soares 1966) [=Piresa villosa]K03b
                  |--Neopachylus Roewer 1913 (see below for synonymy)K03b
                  |    |--*N. bellicosus Roewer 1913K03b
                  |    |--N. herteli Soares & Soares 1945K03b
                  |    |--N. imaguirei Soares & Soares 1947K03b
                  |    |--N. incertus (Mello-Leitão 1935)KM20 (see below for synonymy)
                  |    |--N. mamillosus Roewer 1915K03b
                  |    |--N. marginatus (Mello-Leitão 1931)KM20 (see below for synonymy)
                  |    `--N. taioensis Soares 1966K03b
                  |--Metagyndes Roewer 1913K03b
                  |    |--*M. martensii (Sørensen in Henriksen 1902)K03b, SH20 [=Pachylus martensiiSH20, M. martensis (l. c.)SH20]
                  |    |--M. chilensis Roewer 1943K03b
                  |    |--M. innata Roewer 1929K03b
                  |    |--M. intermedia Roewer 1913K03b
                  |    |--M. laeviscutata Roewer 1943K03b
                  |    |--M. longispina Mello-Leitão 1936K03b
                  |    |--M. pulchellus (Loman 1899)SH20 (see below for synonymy)
                  |    `--M. trifidus Mello-Leitão 1943K03b
                  `--Neopucroliella Roewer 1931 [incl. Thorellidia Mello-Leitão 1931]K03b
                       |--N. pertyi (Thorell 1877) (see below for synonymy)K03b
                       |--N. bruchi Canals 1935 [=Parabalta bruchi]K03b
                       |--N. calamuchitaensis Canals 1943 [=Parabalta calamuchitaensis]K03b
                       |--N. extraordinaria Canals 1943 [=Parabalta extraordinaria]K03b
                       |--N. mesembrina Ringuelet 1961 [=N. borgmeieri mesembrina]K03b
                       |--N. nonoensis Canals 1943 [=Parabalta nonoensis]K03b
                       |--N. ochracea Ringuelet 1963K03b
                       `--N. sanctiludovici Acosta 1993K03b
Gonyleptidae incertae sedis:
  Nanophareus Roewer 1929K03b
    `--*N. palpalis Roewer 1929K03b
  Pachylospeleus Šilhavý 1974K03b [PachylospeleinaeP-R02]
    `--*P. strinatii Šilhavý 1974K03b
  GonyassamiinaeP-R02
  RoeweriinaeKM20
    |--Khazaddum Carvalho, Kury & Hara 2020CKH20
    |    `--*K. inerme (Soares & Soares 1947)CKH20 [=Daguerreia inermisCKH20, Daquerreia inermes (l. c.)K03b]
    `--Roeweria Mello-Leitão 1923K18 (see below for synonymy)
         |--*R. bittencourti Mello-Leitão 1923 (see below for synonymy)K03b
         `--R. virescens (Mello-Leitão 1940)DG09 (see below for synonymy)
  BourguyiinaeM11
  Acanthogonyleptes Mello-Leitão 1922KA20 [incl. Moreira Roewer 1930KA20, Nygoleptes Mello-Leitão 1931K03b]
    |--*A. pulcher Mello-Leitão 1922KA20 (see below for synonymy)
    |--‘Paragonyleptes’ alticola Mello-Leitão 1922 (see below for synonymy)K03b
    |--A. asperulus (Roewer 1930) [=Moreira asperula, Paragonyleptes asperulus]KA20
    |--A. dubius (Mello-Leitão 1932) [=Paragonyleptes dubius, Progonyleptoides dubius]KA20
    |--A. fulvigranulatus (Mello-Leitão 1922)KA20
    |--A. pygoplus (Roewer 1917) [=Paragonyleptes pygoplus]KA20
    `--A. wygodzinskyi (Soares & Soares 1954) [=Metagonyleptes wygodzinskyi]KA20

*Acanthogonyleptes pulcher Mello-Leitão 1922KA20 [=Paragonyleptes pulcherK03b, Sphaerobunus pulcherK03b; incl. Moreira albicirculata Roewer 1943K03b, P. albicirculatusK03b, *Nygoleptes ornatus Mello-Leitão 1931K03b]

Acanthopachylus Roewer 1913 [=Acanthpachylus; incl. Guascaia Mello-Leitão 1935, Heteropachyloidellus Mello-Leitão 1927]K03b

*Acanthopachylus aculeatus (Kirby 1819)K03b, KA-Z11 [=Gonyleptes aculeatusK03b, Acanthpachylus (l. c.) aculeatusK03b, Pachylus aculeatusK03b; incl. ‘Faucheur acanthure’ Duméril 1816-1830K03b, Gonyleptes acanthurusK03b, *Heteropachyloidellus dimorphicus Mello-Leitão 1927K03b, H. marginatus Mello-Leitão 1932K03b, Pachylus mesopotamogalis Weyenbergh 1878K03b, H. paucigranulatus Mello-Leitão 1931K03b, Gonyleptes robustus Holmberg 1876K03b, Pachylus robustusK03b, Heteropachyloidellus robustus Roewer 1943 non Gonyleptes robustus Holmberg 1876K03b, H. crassus (l. c.)K03b, H. serrulatus Mello-Leitão 1931K03b, Pachyloidellus tricalcaratus Roewer 1923K03b, Progyndes tricalcaratusK03b, *Guascaia ypsilonota Mello-Leitão 1935K03b, Guascaia ypsilonata (l. c.)K03b, Guascaya (l. c.) ypsilonotaK03b]

Acanthoprocta Loman 1899 [=Acantoprocta (l. c.), Acantrhoprocta (l. c.); incl. Parapachylus Roewer 1913]K03b

*Acanthoprocta pustulata Loman 1899 [=Acanthroprocta (l. c.) pustulata, Pachylus pustulatus; incl. Parapachylus bispinifrons Roewer 1913, Pachyloides glabrio Loman 1899, *Par. glabrio, Pachylus glabrio]K03b

Acrographinotus Holmgren 1916A02, KA-Z11 [incl. Ctatoproceros Soares & Bauab-Vianna 1973K03b, KA-Z11, Liographinotus Roewer 1957A02]

*Berlaia dissimilis Mello-Leitão 1940 [incl. Echinopachylus brevipes Mello-Leitão in Soares & Soares 1987 (n. n.), Joffilyana nasuta Mello-Leitão in Soares & Soares 1987 (n. n.), Berlaia spinulosa Mello-Leitão 1940]K03b

Carlotta Roewer 1943 [incl. Pseudoneogonyleptoides Soares 1944, Quixaba Mello-Leitão 1944, Quixabella Mello-Leitão 1949, Ullia Roewer 1943]K03b

*Carlotta serratipes Roewer 1943 [incl. *Quixaba atrolutea Mello-Leitão 1944, *Ullia furcata Roewer 1943, Carlotta furcata, *Quixabella ornata Mello-Leitão 1949]K03b

*Ceratoleptes proboscis Soares & Soares 1979 [incl. Ancylorhinoleptes assimilis Mello-Leitão in Kury 2003 (n. n.)]K03b

Cobania Roewer 1913M11, K03b [incl. Crypturocytia Mello-Leitão 1932K03b, Flangeia Mello-Leitão 1933K03b, Liogyndulus Mello-Leitão 1932KP-R20; Cobaniinae]

*Cobania picea (Bertkau 1880) [=Gonyleptes piceus; incl. *Crypturocytia crypturocytia Mello-Leitão 1932, Lyogonyleptoides (l. c.) curvifemur Roewer 1943, Liogonyleptoides curvifemur, *Liogyndulus luteifemur Mello-Leitão 1932]K03b

Deltaspidium asper (Perty 1833) [=Gonyleptes asper, Ampheres asper; incl. *Deltaspidium bresslaui Roewer 1927]K03b

Eubalta planiceps (Gervais 1842)KM20 [=Gonyleptes planicepsKM20, Lycomedes planicepsK03b, Lycomedicus planicepsK03b, Lycomedicus laniceps (l. c.)K03b, Pachylus planicepsK03b, Sadocus planicepsK03b; incl. *Balta meridionalis Sørensen 1902K03b, P-SH20, *Baltanata meridionalisK03b, *Eubalta meridionalisK03b, Gonyleptes modestus Gervais 1847P-SH20, Discocyrtus modestusP-SH20]

Glysterus Roewer 1931KA-Z11 [incl. Angathisoma Roewer 1943K03b, Glysteroides Roewer 1943K03b, Phalacrobunus Roewer 1943K03b, Rooma Goodnight & Goodnight 1942K03b, Triaenoplus Roewer 1943K03b]

Gonyleptellus cancellatus (Roewer 1917) [=Gonyleptes cancellatus, Megapachylus cancellatus, Paragonyleptes cancellatus; incl. *Stephanocranion bimaculatus Mello-Leitão 1931, *Gonyleptellus multimaculatus Roewer 1930, Progonyleptoides pustulosus Mello-Leitão 1935, Stephanocranion serrulatum Mello-Leitão 1940]K03b

*Huadquina huadquinae (Chamberlin 1916) [=Gonoleptes (l. c. for Gonyleptes) huadquinae, Progonyleptoides huadquinae]K03b

*Junicus gerhardi Goodnight & Goodnight 1947 [incl. Acrographinotus luteipalpis Roewer 1957, *Sokkupia olivacea Mello-Leitão 1949, A. olivaceus]K03b

Kuryella melanoacantha (Soares & Soares 1947)O06 [=Paranaleptes melanoacanthusO06, P. xelanoacanthus (l. c.)K03b]

Liogonyleptoides Mello-Leitão 1925 [=Progonyleptoides Mello-Leitão 1922 non Roewer 1917, Lyogonyleptoides (l. c.); incl. Anomaloleptes Mello-Leitão 1935, Pachyleptes Mello-Leitão 1932]K03b

*Liogonyleptoides inermis (Mello-Leitão 1922) [=*Progonyleptoides inermis, Lyogonyleptoides (l. c.) inermis; incl. Liogonyleptoides calcaratus Piza 1936, Progonyleptoides cimex Mello-Leitão 1923, Li. cimex, Lyogonyleptoides (l. c.) cimex, *Anomaloleptes singularis Mello-Leitão 1935]K03b

Mangaratiba acanthoproctus (Soares 1968)KA-Z11 [=*Canopilio acanthoproctusK03, M. acanthoprocta (l. c.)K03]

Megapachylus Roewer 1913 [incl. Gonyleptilus Roewer 1927, Gonyperna Roewer 1930, Pachylometoides Mello-Leitão 1936, Pegada Roewer 1930 non Moore 1892, Pegadana Strand 1942]K03b

*Megapachylus grandis Roewer 1913 [incl. Gonyperna curvispina Mello-Leitão 1933, Paragonyleptes curvispina, *Gonyperna divaricata Roewer 1930, Paragonyleptes divaricatus, *Pachylometoides pugnax Mello-Leitão 1936, *Gonyleptilus umbonatus Roewer 1927, Paragonyleptes umbonatus]K03b

Megapachylus simoni (Roewer 1930) [=Paragonyleptes simoni; incl. *Pegada oedipoda Roewer 1930, *Pegadana oedipoda]K03b

Metagyndes pulchellus (Loman 1899)SH20 [=Gyndes pulchellaSH20, Pachylus pulchellusK03b; incl. Gyndulus curvitibialis Roewer 1943SH20, Progyndes roeweri Soares & Soares 1954SH20, Gyndulus roeweriSH20, Metagyndes roeweriSH20]

Meteusarcoides caudatus (Piza 1940) [=*Cercopachylus caudatus, Uropachylus caudatus; incl. Meteusarcoides marombaensis Soares & Soares 1954]K03b

Neopachylus Roewer 1913 [incl. Gephyropachylus Mello-Leitão 1931, Huralvius Mello-Leitão 1935, Xuraloius (l. c.), Nunduavius Mello-Leitão 1936]K03b

Neopachylus incertus (Mello-Leitão 1935)KM20 [=*Huralvius incertusK03b, Xuraloius (l. c.) incertusK03b; incl. *Nunduavius nebulosus Mello-Leitão 1936K03b, KM20, Neopachylus nebulosusK03b]

Neopachylus marginatus (Mello-Leitão 1931)KM20 [=*Gephyropachylus marginatusK03b; incl. N. serrinha Soares & Soares 1947KM20]

Neopucroliella pertyi (Thorell 1877) [=Ostracidium pertyi, *Thorellidia pertyi; incl. Neopucrolia borgmeieri Mello-Leitão 1923, *Neopucroliella borgmeieri, Neopucroliella borgmeyer (l. c.), Parabalta borgmeieri]K03b

Neosadocus Mello-Leitão 1926KP-R07, K03b [incl. Bunoweyhia Mello-Leitão 1935 (nom. inv.)KA-Z11, Ilhania Mello-Leitão 1936K03b, Mitoperna Roewer 1931K03b, Polybunos Piza 1943K03b]

*Neosadocus bufo (Mello-Leitão 1923) [=Sadocus bufo; incl. Bunoweyhia lata Mello-Leitão 1935, Neosadocus latus, B. minor Mello-Leitão 1935, N. minor, *Polybunos tuberculatus Piza 1943]K03b

Nesopachylus Chamberlin 1925KA-Z11 [=Neopachylus (l. c.) non Roewer 1913K03b; incl. Gorgonasta Roewer 1932K03b]

Ogloblinia Canals 1933 [=Oglobinia (l. c.), Oglobinius (l. c.); incl. Prosampycus Mello-Leitão 1935, Quitete Mello-Leitão 1936]K03b

Ogloblinia argenteopilosa (Mello-Leitão 1935) [=*Prosampycus argenteopilosus, Oglobinia (l. c.) argenteopilosa; incl. *Quitete marginata Mello-Leitão 1936]K03b

*Ogloblinia loretoensis Canals 1933 [=Oglobinia (l. c.) loretoensis; incl. Ogloblinia calcarata Soares & Bauab-Vianna 1970, Ogloblinia dentei Soares 1945, Oglobinia intermedia Soares 1944]K03b

Pachyloidellus Müller 1917K03b, KA-Z11 [=Pachiloidellus (l. c.)K03b; incl. Doelloa Mello-Leitão 1930K03b, Petrocchia Mello-Leitão 1933K03b, Sphaleropachylus Mello-Leitão 1926K03b, Sphalepopachylus (l. c.)K03b, Spraleropachylus (l. c.)K03b]

Pachyloidellus butleri (Thorell 1877) [=Pachylus butleri, Acanthopachylus butleri, Acanthpachylus (l. c.) butleri, *Sphaleropachylus butleri, S. buttleri (l. c.); incl. *Pachyloidellus fuscus Müller 1917, Progyndes fuscus, Pachylus gouldii Weyenbergh 1878, *Petrocchia lesserti Mello-Leitão 1933, Sphaleropachylus lesserti, Pe. marginata Mello-Leitão in Kury 2003 (n. n.)]K03b

Pachyloidellus fulvigranulatus (Mello-Leitão 1930) [=*Doelloa fulvigranulata, Sphaleropachylus fulvigranulatus]K03b

Pachylus chilensis (Gray 1833)KM20 [=Gonoleptes chilensisKM20, Gonyleptes chilensisK03b, Kochipachylus chilensisK03b; incl. Gony. curvipes Guérin-Méneville 1838K03b, *Pachylus granulatus Koch 1839K03b, KM20, *Acanthopachyloides patellaris Piza 1942K03b]

‘Paragonyleptes’ alticola Mello-Leitão 1922 [=*Moreira alticola, Sphaerobunus alticola; incl. Soerensenia transfasciata Mello-Leitão 1931, Paragonyleptes transfasciatus, Soerensia (l. c.) transfasciata]K03b

*Paraluederwaldtia bituberculata (Mello-Leitão 1922) [=Neopucrolia bituberculata, Isopucrolia (l. c.) bituberculata, Luederwaldtia bituberculata]K03b

Parampheres Roewer 1913KP-R07, K03b [incl. Callampheres Roewer 1931K03b, Cezarella Mello-Leitão 1932K03b, Pertyana Mello-Leitão 1927K03b]

Parampheres bimaculatus (Mello-Leitão 1932) [=*Cezarella bimaculata, Cearinus corniger bimaculatus, Ilhaia bimaculata]K03b

Parampheres ronae (Mello-Leitão 1927) [=*Pertyana ronae, Per. ronnae (l. c.); incl. Penygorna bimaculata Mello-Leitão 1937 non Cezarella bimaculata Mello-Leitão 1932, Arleius bimaculatus, Ilhaia bimaculata non (Mello-Leitão 1932), Parampheres bimaculatus Roewer 1943 nec Cezarella bimaculata Mello-Leitão 1932 nec Penygorna bimaculata Mello-Leitão 1937, Pa. nigrimanus Mello-Leitão 1933, Ilhaia ringueleti Capocasale 1973]K03b

Parapachyloides Roewer 1913 [incl. Apophysigerus Canals 1935, Goyazella Mello-Leitão 1931, Tabatinguera Mello-Leitão 1935]K03b

Parapachyloides armatus (Mello-Leitão 1931) [=*Goyazella armata; incl. *Tabatinguera insignis Mello-Leitão 1935, Parapachyloides insignis]K03b

*Parapucrolia ocellata Roewer 1917 [incl. *Goytacazia pulchra Mello-Leitão 1942, Goitacazia (l. c.) pulchra, Goytacazio (l. c.) pulchra]K03b

Pygophalangodus Mello-Leitão 1931K03b [incl. Bacigalupo Mello-Leitão 1933K03b, Canestrinia Mello-Leitão 1931 nec Berlese 1881 nec Mégnin & Trouessart 1884K03b, Melloinia Thor 1933K03b, Melloinio (l. c.)KA-Z11, Melloleitaoella Strand 1932K03b]

Pygophalangodus canalsi (Mello-Leitão 1931) [=*Canestrinia canalsi, *Melloinia canalsi, *Melloleitaoella canalsi; incl. *Bacigalupo tenax Mello-Leitão 1933]K03b

Roeweria Mello-Leitão 1923K18 [incl. Jupuvura Mello-Leitão 1940K03b, Lyopachylus Mello-Leitão 1926K03b, Liopachylus (l. c.)K03b, Paranaincola Soares 1943K03b]

*Roeweria bittencourti Mello-Leitão 1923 [=Discocyrtoides bittencourti; incl. D. catarinae Soares 1946, *Lyopachylus mitobatoides Mello-Leitão 1923]K03b

Roeweria virescens (Mello-Leitão 1940)DG09 [=*Jupuvura virescensK03b, Discocyrtoides virescensK03b; incl. Discocyrtus fortis Soares 1945K03b, *Paranaincola mourei Soares 1943K03b, Discocyrtus moureiK03b, Lyopachylus moureiK03b]

Schenkelibunus Strand 1932K03b, KA-Z11 [incl. Friburgoia Mello-Leitão 1932KA-Z11, Hanseniella Mello-Leitão 1927 non Bagnall 1913K03b, Ziltaia Mello-Leitão 1936K03b, Siltaya (l. c.)K03b]

Schenkelibunus impar (Mello-Leitão 1932)KA-Z11 [=*Friburgoia imparK03b, Hanseniella imparK03b; incl. *Ziltaia nigrifemur Mello-Leitão 1936K03b, Ilhaia nigrifemurK03b]

Sphaerobunus Roewer 1917DG09, K03b [incl. Adelphobunus Mello-Leitão 1935K03b, Bocaina Piza 1943K03b, Caldasiella Mello-Leitão 1931K03b, Caxambusia Mello-Leitão 1935K03b, Costalimaiella Mello-Leitão 1931K03b, Costalimarella (l. c.)K03b, Diplocaldasius Mello-Leitão 1934K03b, Kaingangoides Piza 1938K03b, Leptogonys Mello-Leitão 1931K03b, Monocerodynus Mello-Leitão 1940K03b, Paranastus Roewer 1943K03b, Soerensenia Mello-Leitão 1926K03b, Soerensia (l. c.)K03b, Xenoleptes Mello-Leitão 1942K03b]

Sphaerobunus editus (Roewer 1943) [=*Paranastus editus, Paragonyleptes editus; incl. Paragonyleptes serranus Soares 1945]K03b

Sphaerobunus fallax (Mello-Leitão 1932) [=Metagonyleptes fallax, Paragonyleptes fallax; incl. *Leptogonys marmoratus Mello-Leitão 1932 non Progonyleptes marmoratus Mello-Leitão 1940, Pa. marmoratus]K03b

Sphaerobunus fulvigranulatus (Mello-Leitão 1922) [=Paragonyleptes fulvigranulatus, *Soerensenia fulvigranulata; incl. *Costalimaiella conspicua Mello-Leitão 1931, Gonyperna incus Piza 1943, *Caldasiella nigra Mello-Leitão 1931, Paragonyleptes niger, P. nigrus, *Diplocaldasius pallidus Mello-Leitão 1934, *Kaingangoides pardalis Piza 1938, Paragonyleptes pardalis, Caldasiella pustulosa Mello-Leitão 1939, P. pustulosus, Metagonyleptes pygoplus Mello-Leitão 1932, P. triacanthus Mello-Leitão 1922]K03b

Sphaerobunus pictus (Piza 1942) [=Paragonyleptes pictus; incl. Caldasiella marmorata Mello-Leitão 1933 (nom. inval.) non Leptogonys marmoratus Mello-Leitão 1932]K03b

Sphaerobunus singularis (Mello-Leitão 1935) [=*Adelphobunus singularis; incl. *Bocaina marmorata Piza 1943 nec Leptogonys marmoratus Mello-Leitão 1932 nec Caldasiella marmorata Mello-Leitão 1933 nec Progonyleptes marmoratus Mello-Leitão 1940, Adelphobunus pulcher Mello-Leitão 1935 non Acanthogonyleptes pulcher Mello-Leitão 1935, *Caxambusia variegata Mello-Leitão 1935]K03b

Sphaerobunus soaresi (Mello-Leitão 1944) [=Paragonyleptes soaresi; incl. Caldasiella anomala Soares 1942 (nom. inval.), Paragonyleptes anomalus non Mello-Leitão 1922]K03b

Sphaerobunus variolosus (Mello-Leitão 1940) [=*Monocerodynus variolosus; incl. *Xenoleptes nigrimanus Mello-Leitão 1942]K03b

Tingomaria Mello-Leitão in Mello-Leitão & Feio 1949 1949K03b, KM20 [incl. Goyazelloides Soares & Bauab-Vianna 1972K03b]

*Tingomaria hydrophila Mello-Leitão in Mello-Leitão 1949KM20 [=*Goyazelloides peruvicus Soares & Bauab-Vianna 1972K03b]

Triglochinura Mello-Leitão 1924 [incl. Barbiellinia Mello-Leitão 1944 non Bezzi 1922, Itaoca Mello-Leitão 1935, Piresa Roewer 1927, Tayoca Mello-Leitão 1937]K03b

*Triglochinura curvispina Mello-Leitão 1924K03b [incl. *Piresa armata Roewer 1927K03b, T. armataA-NM06, *Barbiellinia hirsuta Mello-Leitão 1944K03b, Piresa hirsutaK03b, *Itaoca melanacantha Mello-Leitão 1935K03b, P. melanacanthaK03b, P. sanctipauli Soares 1945K03b]

*Uracantholeptes anomalus (Mello-Leitão 1922) [=Paragonyleptes anomalus; incl. *Gonypernoides fragilis Mello-Leitão 1932]K03b

Uropachylus Mello-Leitão 1922 [=Japyra Mello-Leitão 1934, Japira (l. c.), Ypiranga Mello-Leitão 1922, Ipiranga (l. c.)]K03b

Uropachylus ypiranga (Mello-Leitão 1922) [=*Ypiranga ypiranga; incl. Cercopachylus fragilis Piza 1943, *Japyra regularis Mello-Leitão 1934]K03b

*Type species of generic name indicated

References

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[A-NM06] Almeida-Neto, M., G. Machado, R. Pinto-da-Rocha & A. A. Giaretta. 2006. Harvestman (Arachnida: Opiliones) species distribution along three Neotropical elevational gradients: an alternative rescue effect to explain Rapoport’s rule? Journal of Biogeography 33: 361–375.

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