Gymnobela engonia, copyright Smithsonian National Museum of Natural History.

Belongs within: Raphitomidae.

Gymnobela is a genus of conoids with a short and broad, often rather solid shell and lacking an operculum (Bouchet & Waren 1980).

<==Gymnobela Verrill 1884 [incl. Majox Nordsieck 1968, Watsonaria Nordsieck 1968]BK11
    |--G. (Gymnobela)P66
    |    |--*G. (G.) engonia Verrill 1884BW80
    |    |--G. (G.) agonia (Dall 1889) [=Pleurotomella (G.) agonia]P66
    |    |    |--G. a. agoniaP66
    |    |    `--G. a. altina (Dall 1908) [=Pleurotomella (Gymnobela) altina]P66
    |    |--G. (G.) egregia (Dall 1908) [=Pleurotomella (G.) egregia]P66
    |    |--G. (G.) isogonia (Dall 1908) [=Pleurotomella (G.) isogonia]P66
    |    `--G. (G.) xylona (Dall 1908) [=Pleurotomella (G.) xylona]P66
    `--G. (Theta Clarke 1959)BK11
         |--‘Pleurotomella’ (*T.) lyronuclea Clarke 1959 [=Pleurotomoides lyronuclea]BW80
         |--‘Theta’ chariessa (Watson 1881)BW80 (see below for synonymy)
         `--‘Theta’ vayssierei (Dautzenberg 1925) (see below for synonymy)BW80
Gymnobela incertae sedis:
  G. abyssorum (Locard 1897) [=Bela abyssorum]BW80
  G. agassizi (Verrill & Smith in Verrill 1880) (see below for synonymy)BW80
    |--G. a. agassiziP66
    `--‘Pleurotomella’ a. mexicana Dall 1889P66
  G. aquilarum (Watson 1881)BW80 (see below for synonymy)
  G. augusta Thiele 1925P66
  G. bairdi (Verrill & Smith 1884) [=Pleurotomella bairdi, *Majox bairdi]BW80
  G. blakeana (Dall 1881) [=Pleurotoma (Bela) blakeana]P66
    |--G. b. blakeanaP66
    `--G. b. agria Dall 1889P66
  G. brevis (Bellardi 1847) [=Pleurotoma brevis]BW80
  G. camerunensis Thiele 1925P66
  G. chrysopelex (Barnard 1963) [=Typhlosyrinx chrysopelex]BS01
  G. chyta (Watson 1881)BW80 [=Pleurotoma (Defrancia) chytaBW80, Clathurella chytaBW80, Pleurotomella chytaP66]
  G. clara Thiele 1925P66
  G. dautzenbergi (Knudsen 1952) [=Cythara dautzenbergi]RO-SF98
  G. emertoni (Verrill & Smith 1884)BW80 (see below for synonymy)
  G. extensa (Dall 1881) (see below for synonymy)P66
  G. frielei (Verrill 1885)BW80 (see below for synonymy)
  G. fulvotincta (Dautzenberg & Fischer 1896)BW80 [=Pleurotoma fulvotinctaBW80, Pleurotomella fulvotinctaP66]
  G. grundifera Dall 1927P66
  G. homeotata (Watson 1886)BW80 [=Clathurella homeotataBW80, Pleurotomella homaeotataP66]
  G. illicita Dall 1927P66
  G. imitator Dall 1927P66
  G. lamyi (Dautzenbereg 1925) [=Pleurotoma (Pleurotomella) lamyi]BW80
  G. lanceata Dall 1927P66
  G. leptoglypta (Dautzenberg & Fischer 1896)BW80 (see below for synonymy)
  G. phyxanor (Watson 1886)BW80 [=Clathurella phyxanorBW80, Pleurotomella phyxanorP66]
  G. pyrrhogramma (Dautzenberg & Fischer 1896)BW80 (see below for synonymy)
  G. ryukyuensis (MacNeil 1961)BS01
  G. subaraneosa (Dautzenberg & Fischer 1896)BW80 [=Pleurotoma subaraneosaBW80, Pleurotomella subaraneosaP66]
  G. watsoni (Dautzenberg 1889) [=Clathurella watsoni, Pleurotomella watsoni, *Watsonaria watsoni]BW80
  G. yoshidai Kuroda & Habe in Habe 1962BK11

*Type species of generic name indicated


[BK11] Bouchet, P., Y. I. Kantor, A. Sysoev & N. Puillandre. 2011. A new operational classification of the Conoidea (Gastropoda). Journal of Molluscan Studies 77: 273–308.

[BW80] Bouchet, P., & A. Waren. 1980. Revision of the north-east Atlantic bathyal and abyssal Turridae (Mollusca, Gastropoda). Journal of Molluscan Studies Supplement 8: 1–119.

[BS01] Bouchet, P., & A. Sysoev. 2001. Typhlosyrinx-like tropical deep-water turriform gastropods (Mollusca, Gastropoda, Conoidea). Journal of Natural History 35: 1693–1715.

[P66] Powell, A. W. B. 1966. The molluscan families Speightiidae and Turridae: An evaluation of the valid taxa, both Recent and fossil, with lists of characteristic species. Bulletin of the Auckland Institute and Museum 5: 1–184, pls 1–23.

[RO-SF98] Rolán, E., J. Otero-Schmitt & F. Fernandes. 1998. The family Turridae s.l. (Mollusca, Gastropoda) in Angola (West Africa), 1. Subfamily Daphnellinae. Iberus 16 (1): 95–118.

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