Gymnomyzinae

Placopsidella grandis, from the National Museum of Natural History.

Belongs within: Ephydroidea.

The Gymnomyzinae are a group of shore flies with typically pectinate aristae and small prescutellar acrostichal setae. Representatives include the cosmopolitan predatory genus Ochthera in which the fore legs are raptorial with the femur enlarged and the tibia bearing a large apical spur. Members of the tribe Gastropsini have a trigonal face, and abdominal tergites 3 and 4 enlarged and rugose or pitted (Mathis 2010).

Characters (from Mathis 2010): Arista pectinate; reclinate fronto-orbital seta usually inserted slightly anterior to larger, proclinate fronto-orbital seta; median facial area and lower facial margin without setae; facial setae inserted in more or less vertical series, parallel with parafacial setae; prescutellar acrostichal setae small (about one-half to two-thirds length of posterior dorsocentral seta), inserted close together (distance between about one-half that between either prescutellar and posterior dorsocentral seta on the same side), behind or aligned with intra-alar seta.

<==GymnomyzinaeM10
    |--Ochthera Latreille 1802L02 [OchtheriniM10]
    |    |--O. manicata [=Musca manicata]L02
    |    `--O. mantisM10 [=Musca mantisL02]
    |--Hecamede [Hecamedini]M10
    |    |--H. albicansM10
    |    `--H. brasiliensisM10
    |--AtissiniM10
    |    |--Ptilomyia mabelaeM10
    |    `--AtissaM10
    |         |--A. atlanticaM10
    |         `--A. pygmaeaWMV87
    |--GastropsiniM10
    |    |--BeckeriellaM10
    |    `--GastropsM10
    |         |--G. nebulosusM10
    |         `--G. nigerM10
    |--DiscocerininiM10
    |    |--Discocerina obscurellaM10
    |    |--GymnoclasiopaM10
    |    |--Pectinifer aeneusM10
    |    |--DitrichophoraM10
    |    |--Polytrichophora agensM10
    |    |--Hydrochasma leucoproctumM10
    |    |--LamproclasiopaM10
    |    `--Orasiopa meraM10
    |--LipochaetiniM10
    |    |--Glenanthe interiorM10
    |    |--Paraglenanthe bahamensisM10, WMV87
    |    |--LipochaetaM10
    |    |    |--L. ranicaM10
    |    |    `--L. slossonaeM10
    |    |--DiphuiaM10
    |    |    |--D. anomalaM10
    |    |    `--D. nitidaM10
    |    `--AllotrichomaM10
    |         |--A. (Pseudohecamede) abdominaleM10 [=Pseudohecamede abdominalisWMV87]
    |         `--A. simplexM10
    `--GymnomyziniM10
         |--Cerometopum lacunosumM10
         |--AthyroglossaM10
         |    |--A. glabraM10
         |    |--A. glaphyropusM10
         |    `--A. granulosaM10
         |--PlacopsidellaM10
         |    |--P. cynocephalaM10
         |    |--P. grandisM10
         |    `--P. marquesanaM10
         `--MosillusM10
              |--M. arcuatusR26
              |--M. cellariaR26 [=Micropeza (Mosillus) cellarisG20]
              |--M. erythrophthalmaR26
              |--M. stegmaieriM10
              `--M. tibialisM10

*Type species of generic name indicated

References

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[M10] Mathis, W. N. 2010. Ephydridae (shore flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 2 pp. 1211–1233. NRC Research Press: Ottawa.

[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.

[WMV87] Wirth, W. W., W. N. Mathis & J. R. Vockeroth. 1987. Ephydridae. In: McAlpine, J. F. (ed.) Manual of Nearctic Diptera vol. 2 pp. 1027–1047. Research Branch, Agriculture Canada.

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