Hadrinae

Rhagada convicta, copyright G. & Ph. Poppe.

Belongs within: Helicoidei.

The Hadrinae are a group of land snails found in Australia and southeast Asia united by molecular analysis. Members of this clade either lack a head wart or have the head wart (positioned between and just behind the ommatophores) exposed and not eversible (Solem 1997).

<==Hadrinae [Hadridae, Sinumeloninae]BR17
|--Micromelon Solem 1992S97
|--Lacustrelix Iredale 1937S97
|--Basedowena Iredale 1937S97
|--Tatemelon Solem 1993S97
|--Eximiorhagada Iredale 1933S97
|--Montanomelon Solem 1993S97
|--Chloritis Beck 1837BR17, BR05 [Chloritidae]
| `--*C. ungulina (Linnaeus 1758) [=Helix ungulina]BR17
|--Calycia Adams 1865BR17, BR05 [Calyciidae]
| `--*C. crystallina (Reeve 1848) [=Bulimus crystallinus]BR17
|--Planispira Beck 1837BR17, BR05 [Planispiridae]
| `--*P. zonaria (Linnaeus 1767) [=Helix zonaria]BR17
|--Minimelon Solem 1993S97
| `--M. colmani Solem 1993S97
|--Falspleuroxia Solem 1997S97
| `--*F. overlanderensis Solem 1997S97
|--Promonturconchum Solem 1997S97
| `--*P. superbum Solem 1997S97
|--Caperantrum Solem 1997S97
| `--*C. polygyrum Solem 1997S97
|--Cristovala Clench 1958BR17, BR05 [Cristovalinae]
| `--*C. tricolor (Pfeiffer 1849) [=Helix tricolor]BR17
|--Granulomelon Iredale 1933S97
| |--G. arcigerens (Tate 1894)S97
| `--G. gilleni Solem 1993S97
|--Papuina Martens 1860BR17, BR05 [incl. Zetomina Iredale 1941H64; Papuinidae, Papuininae]
| |--*P. lituus (Lesson 1831) [=Helix lituus]BR17
| |--P. exsultansH64
| |--P. ferussaciH64
| `--P. steursiana (Shuttleworth 1852) (see below for synonymy)H64
|--Strepsitaurus Solem 1997S97
| |--*S. rugus (Cotton 1953) [=Pleuroxia ruga]S97
| |--S. cardabius Solem 1997S97
| |--S. milyeringus Solem 1997S97
| |--S. ningaloo Solem 1997S97
| `--S. williami Solem 1997S97
|--Plectorhagada Iredale 1933 [incl. Idamera Iredale 1939]S97
| |--*P. plectilis (Benson 1853)S97 (see below for synonymy)
| |--P. carcharias (Pfeiffer 1864)W08 (see below for synonymy)
| |--P. gascoynensis (Smith 1894) [=Helix (Trachia) gascoynensis, Planispira gascoynensis, Pleuroxia gascoynensis]S97
| |--P. meilgana Solem 1997S97
| |--P. rovina Iredale 1939 [=P. (*Idamera) rovina]S97
| `--P. scolythra Solem 1997S97
|--Hadra Martens 1860BR17 [incl. Micardista Iredale 1933SS10]
| |--*H. bipartita (Férussac 1823) [=Helix bipartita]SS10
| |--H. barneyi (Cox 1873)SS10
| |--H. bartschi (Marshall 1927) (see below for synonymy)SS10
| |--H. brunodavidi Stanisic in Stanisic, Shea et al. 2010SS10
| |--H. funiculata (Reeve 1854)SS10
| |--H. semicastanea (Pfeiffer 1849) (see below for synonymy)SS10
| |--H. webbi (Pilsbry 1900) [incl. H. webbi incallida Iredale 1933]SS10
| `--H. wilsoni Solem 1979S97
|--Xanthomelon Martens 1860 [Xanthomelontidae, Xanthomelontinae]BR17
| |--*X. pomum (Pfeiffer 1842) [=Helix pomum]SS10
| |--X. distractum Iredale 1938SS10
| |--X. jannellei (Le Guillou 1842) (see below for synonymy)SS10
| |--X. magnidicum Iredale 1938 [=X. pachystylum magnidicum]SS10
| |--X. obliquirugosa (Smith 1894) [=Globorhagada obliquirugosa]S97
| |--X. pachystylum (Pfeiffer 1845) (see below for synonymy)SS10
| |--X. prudhoensis (Smith 1894) [=Globorhagada prudhoensis]S97
| `--X. saginatum Iredale 1938 [=X. pachystylum saginatum]SS10
|--Pleuroxia Ancey 1887 (see below for synonymy)S97
| |--*P. cyrtopleura (Pfeiffer 1862) [=Helix cyrtopleura]S97
| |--P. abstans Iredale 1939 [=P. (*Gantomia) abstans]S97
| |--P. adcockiana (Bednall 1894)S97
| |--P. bethana Solem 1997S97
| |--P. carmeena Solem 1993S97
| |--P. elfina Iredale 1939 [=P. polypleura elfina]S97
| |--P. everardenis (Bednall 1892)S97
| |--P. hinsbyi (Gude 1916)S97
| |--P. italowiana Solem 1992S97
| |--P. oligopleura (Tate 1894) (see below for synonymy)S97
| |--P. phillipsiana (Angas 1873)S97
| |--P. polypleura (Tate 1899) [=Helix polypleura, Angasella polypleura; incl. P. commenta Iredale 1939]S97
| `--P. radiata (Hedley 1905)S97
|--Sinumelon Iredale 1930BR05 [incl. Notobadistes Cotton & Godfrey 1932S97]
| |--*S. nullaboricum (Tate 1879)BR17 (see below for synonymy)
| |--S. amatensisBRW98
| |--S. bednalliBRW98
| |--‘Helix’ bitaeniata Cox 1868 [=*Notobadistes bitaeniatus]S97
| |--S. dulcensisBRW98
| |--S. expositumBRW98
| |--S. gawleri Solem 1992S97
| |--S. gillensisBRW98
| |--S. hullanumBRW98
| |--S. jimberlanensis Solem 1997S97
| |--S. kalgum Iredale 1939 [incl. S. lennum Iredale 1939, S. lennum mutuum Iredale 1939]S97
| |--S. musgraveiBRW98
| |--S. pedasumBRW98
| |--S. perinflatumBRW98
| |--S. pumilioBRW98
| |--S. tarcoolanum Solem 1992S97
| |--S. vagente Iredale 1939S97
| `--S. wilpenensis (Tate 1894)S97
`--Rhagada Martens 1860BR17 [incl. Bellrhagada Iredale 1938S97, Tumegada Iredale 1939S97; Rhagadidae, Rhagadinae]
| i. s.: *R. reinga (Gray 1846)BR17 [=Helix reingaBR17, Thersites (*Rhagada) reingaS97]
| R. angulata Solem 1997S97
| R. basedowana Iredale 1939S97
| R. biggeanaGK12
| R. bulgana Solem 1997S97
| R. crystalla Solem 1985S97
| R. cygna Solem 1997S97
| R. dampierana Solem 1997S97
| R. dominicaGYD14
| R. dringi (Pfeiffer 1846) [=Helix dringi, Bellrhagada dringi]S97
| R. elachystoma (von Martens 1878) [=Helix elachystoma, Thersites (Rhagada) elachystoma]S97
| R. felicitasGK12
| R. globosa Solem 1997S97
| R. harti Solem 1985S97
| R. herbertiJHF06
| R. intermedia Solem 1997S97
| R. kessneriGK12
| R. minima Solem 1997S97
| R. primigenaGK12
| R. radleyi Preston 1908S97
| R. richardsonii (Smith 1874) (see below for synonymy)S97
| R. sheaeiGK12
| R. supracostulata Schepman 1892S97
| R. tescorum (Benson 1853) [=Helix tescorum]S97
| R. torulus (Férussac 1819) [=Helix torulus, Thersites (Rhagada) torulus]S97
|--+--R. barrowensis Johnson, Stankowski et al. 2013JS13
| `--+--R. convicta (Cox 1870)JS13, S97 [=Helix convictaS97, R. (*Tumegada) convictaS97, Thersites (Rhagada) convictaS97]
| `--R. pilbarana Solem 1997JS13, S97
`--+--R. capensis Solem 1997JS13, S97
`--+--R. perprima Iredale 1939JS13, S97 [=R. convicta perprimaS97]
`--R. plicata Preston 1914JS13, S97 [=*Bellrhagada plicataS97]

Hadra bartschi (Marshall 1927) [incl. Thersites (Hadra) bartschi cepa Marshall 1927, T. (H.) dalli Marshall 1927, T. (H.) bartschi diva Marshall 1927, T. (H.) bartschi elfa Marshall 1927, T. (H.) bartschi fama Marshall 1927, T. (H.) bartschi mobiagensis Marshall 1927, T. (H.) bartschi murrayensis Marshall 1927, T. (H.) bartschi nesia Marshall 1927, T. (H.) bartschi nura Marshall 1927, T. (H.) bartschi oma Marshall 1927, T. (H.) bartschi paulensis Marshall 1927, T. (H.) bastschi quaesita Iredale 1937, T. (H.) waltoni Marshall 1927, T. (H.) bartschi yamensis Marshall 1927]SS10

Hadra semicastanea (Pfeiffer 1849) [incl. Thersites (Hadra) semicastanea alma Marshall 1927, Hadra blighi Iredale 1937, T. (H.) lizardensis Marshall 1927, T. (H.) lizardensis rada Marshall 1927, T. (H.) lizardensis suma Marshall 1927]SS10

Papuina steursiana (Shuttleworth 1852) [=Helix steursiana; incl. P. canefriana Kobelt 1894, P. hedleyi var. concolor Smith 1897, P. hedleyi Smith 1892, *Zetomina hedleyi]H64

Plectorhagada carcharias (Pfeiffer 1864)W08 [=Helix carchariasS97, H. (Dorcasia) carchariasS97, Thersites (Rhagada) carchariasS97]

*Plectorhagada plectilis (Benson 1853)S97 [=Helix plectilisS97, Amplirhagada plectilisW08, H. (Dorcasia) plectilisS97, Thersites (Rhagada) plectilisS97; incl. H. paleata Reeve 1854S97]

Pleuroxia Ancey 1887 [incl. Angasella Adams 1864 non Angasiella Crosse 1864, Angasietta Iredale 1939, Gantomia Iredale 1939]S97

Pleuroxia oligopleura (Tate 1894) [=Hadra oligopleura, Angasella oligopleura; incl. A. lemani Gude 1916, Pleuroxia lemani, P. oligopleura numba Iredale 1939]S97

Rhagada richardsonii (Smith 1874) [=Helix richardsonii, Thersites (Rhagada) richardsonii; incl. Rhagada convicta strella Iredale 1939, R. convicta tambra Iredale 1939]S97

*Sinumelon nullaboricum (Tate 1879)BR17 [=Helix nullaboricaBR17, H. (Xanthomelon) nullarboricaS97, Notobadistes nullarboricaS97, Thersites (Badistes) nullarboricaS97; incl. Sinumelon datum Iredale 1939S97]

Xanthomelon jannellei (Le Guillou 1842) [=Helix jannellei; incl. H. pachystyla daemeli von Martens 1869, H. pachystyloides Cox 1868]SS10

Xanthomelon pachystylum (Pfeiffer 1845) [incl. X. pachystylum noscitum Iredale 1938, Thersites pachystyla subfuscozonata Cox 1894]SS10

*Type species of generic name indicated

References

[BRW98] Beesley, P. L., G. J. B. Ross & A. Wells (eds) 1998. Fauna of Australia vol. 5. Mollusca: The Southern Synthesis. Part A. Australian Biological Resources Study: Canberra.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[GK12] Gibson, L. A., & F. Köhler. 2012. Determinants of species richness and similarity of species composition of land snail communities on Kimberley islands. Records of the Western Australian Museum Supplement 81: 41–66.

[GYD14] Gibson, L. A., S. Yates & P. Doughty (eds) 2012–2014. Biodiversity Values on Selected Kimberley Islands, Australia. Records of the Western Australian Museum Supplement 81.

[H64] Henrard, J. B. 1964. Identification of Helix steursiana. Nautilus 77 (4): 112–116.

[JHF06] Johnson, M. S., Z. R. Hamilton & J. Fitzpatrick. 2006. Genetic diversity of Rhagada land snails on Barrow Island. Journal of the Royal Society of Western Australia 89 (2): 45–50.

[JS13] Johnson, M. S., S. Stankowski, C. S. Whisson, R. J. Teale & Z. R. Hamilton. 2013. Camaenid land snails on Barrow Island: distributions, molecular phylogenetics and taxonomic revision. Records of the Western Australian Museum Supplement 83: 159–171.

[S97] Solem, A. 1997. Camaenid land snails from Western and central Australia (Mollusca: Pulmonata: Camaenidae). VII. Taxa from Dampierland through the Nullarbor. Records of the Western Australian Museum Supplement 50: 1461–1906.

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

[W08] Wilson, B. 2008. Terrestrial gastropods of Faure Island, Shark Bay, Western Australia. Records of the Western Australian Museum Supplement 75: 21–24.

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