Ventral (left) and dorsal views of Halarachne halichoeri, from Reckendorf et al. (2019).

Belongs within: Gamasina.

The Halarachnidae are a group of mesostigmatan mites parasitic in the respiratory tracts of mammals. They usually have the epigynal shield absent or rudimentary whereas the sternal shield is usually present, and stigmata are in a ventral or lateroventral position (Lindquist et al. 2009).

<==Halarachnidae [Halarachninae]LKW09
    |--Zumptiella bakeriLKW09
    |--Pneumonyssoides Fain 1955H98
    |    `--P. caninum (Chandler & Ruhe 1940) [=Pneumonyssus caninum]H98
    |--Orthohalarachne Newell 1947H98
    |    |--O. attenuata (Banks 1910) [=Halarachne attenuata; incl. H. reflexa Tubb 1937, Orthohalarachne reflexa]H98
    |    |--O. diminuata (Doetschmann 1944) [=Halarachne diminuata]H98
    |    `--O. letalis Popp 1961P61
    |--Pneumonyssus Banks 1901 [incl. Pneumotuber Landois & Hoepke 1914]E57
    |    |--*P. simicola Banks 1901H98
    |    `--P. capricorni Domrow 1974H98
    |--Raillietia Trouessart 1902LKW09, H98 [Raillietiidae, Raillietiinae]
    |    |--R. auris (Leidy 1872) [=Gamasus auris]H98
    |    |--R. australis Domrow 1961H98
    |    `--R. caprae Quintero et al. 1980 [incl. R. manfredi Domrow 1981]H98
    `--Halarachne Allman 1847 [incl. Rhinacarus Nehring 1884, Rhinixodes Nehring 1884]ET79
         |--*H. halichoeri Allman 1884ET79
         |--H. americanaLKW09
         `--H. otariaeY91

*Type species of generic name indicated


[E57] Evans, G. O. 1957. An introduction to the British Mesostigmata (Acarina) with keys to families and genera. Journal of the Linnean Society of London, Zoology 43: 203–259.

[ET79] Evans, G. O., & W. M. Till. 1979. Mesostigmatic mites of Britain and Ireland (Chelicerata: Acari-Parasitiformes). An introduction to their external morphology and classification. Transactions of the Zoological Society of London 35: 139–270.

[H98] Halliday, R. B. 1998. Mites of Australia: A checklist and bibliography. CSIRO Publishing: Collingwood.

[LKW09] Lindquist, E. E., G. W. Krantz & D. E. Walter. 2009. Order Mesostigmata. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 124–232. Texas Tech University Press.

[P61] Popp, E. 1961. Orthohalarachne letalis n. sp. (Halarachnidae Oudemans 1906—Mesostigmata), eine neue Milbe bei Zalophus californianus Less. (Otariidae). Acarologia 3 (3): 265–273.

[Y91] Yastrebtsov, A. V. 1991. Features of the embryonic development of gamasid mites (Parasitiformes, Gamasida). In: Dusbábek, F., & V. Bukva (eds) Modern Acarology: Proceedings of the VIII International Congress of Acarology, held in České Budĕjovice, Czechoslovakia, 6–11 August 1990 vol. 2 pp. 115–119. SPB Academic Publishing: The Hague.

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