Madras harp Harpa davidus, copyright H. Zell.

Belongs within: Neogastropoda.

The Harpidae, harp shells, are a group of predatory gastropods with usually colourfully patterned, polished shells bearing strong axial ribs.

Characters (from Wilson & Gillett 1971): Shell usually shiny, strikingly coloured and patterned, sculptured with strong axial ribs more highly polished than remainder of shell. Anterior siphon canal a deep, wide notch; columella lacking folds; operculum absent. Respiratory siphon and eye stalks long; foos broad with front end greatly expanded laterally to form wide, flat, muscular organ.

|--Eoharpa sinuosa Stephenson 1955TTE93
|--Eocithara Fischer 1883WG71
| `--*E. muticaW94
|--Cryptochorda Mörch 1858 [Cryptochordidae, Cryptochordinae]BR05
| `--*C. stromboides (Hermann 1781)BR17 [=Buccinum stromboidesBR17, Buccinopsis stromboidesCC95]
|--Austroharpa Finlay 1931WG71
| |--*A. (Austroharpa) pulligera [=Harpa pulligera]W94
| `--A. (Palamharpa Iredale 1931)W94
| |--A. (*P.) exquisita (Iredale 1931) [=*Palamharpa exquisita]W94
| |--A. (P.) loisae Rehder 1973W94
| |--A. (P.) punctata (Verco 1896)W94
| `--A. (P.) wilsoni Rehder 1973W94
|--Harpa Röding 1798 [Harpina]BR05
| |--*H. harpa (Linnaeus 1758)BR17 [=Buccinum harpaBR17; incl. H. nobilis Röding 1798WG71]
| |--H. amouretta Röding 1798 [incl. H. minor Lamarck 1822]W94
| |--H. articularis Lamarck 1822WG71
| |--H. crenata [incl. H. rivoliana]C64
| |--H. davidus Röding 1798WG71
| |--H. goodwini Rehder 1993BC01
| |--H. gracilis Broderip & Sowerby 1829WG71
| |--H. kajiyamai Habe & Kosuge 1970PK11
| |--H. major Röding 1798WG71
| |--H. neozelanica [=Eocithara neozelanica]A27
| `--H. ventricosaH04
`--Morum Röding 1798 [Moruminae]BR05
| i. s.: M. vicdani Emerson 1995BC01
|--M. (Morum)BR17
| |--*M. (M.) purpureum Röding 1798BR17
| `--M. (M.) oniscus (Linnaeus 1767) [=Strombus oniscus]C80
|--M. (Herculea Adams & Adams 1858 non Fr. 1823 [ICBN])W94
| `--M. (*H.) ponderosum (Hanley 1858) [=Oniscia ponderosa]W94
`--M. (Oniscidia Mörch 1852)BC01 (see below for synonymy)
|--M. (*O.) cancellatumW94 [=Oniscia cancellatumW94, Cassidaria cancellataR42]
|--M. (O.) bruuni Powell 1958 [incl. *Pulchroniscia delecta Garrard 1961]W94
|--M. (O.) exquisitum (Adams & Reeve 1848) [=Oniscia exquisita]C80
|--M. (O.) grande (Adams 1855) [=*Cancellomorum grande, *Onimusiro grandis]W94
|--M. (O.) harpaformis Powell & Bartrum 1929PB29
|--M. (O.) joelgreenei Emerson 1981BC01
|--M. (O.) macdonaldi Emerson 1981BC01
|--M. (O.) matthewsi Emerson 1967 [=M. (Cancellomorum) matthewsi]BC01
|--M. (O.) ninomiyai Emerson 1986BC01
`--M. (O.) veleroae Emerson 1968 [=M. (Cancellomorum) veleroae]BC01

Morum (Oniscidia Mörch 1852)BC01 [incl. Cancellomorum Emerson & Old 1963BC01, Onimusiro Kira in Kuroda, Habe & Oyama 1971W94, Pulchroniscia Garrard 1961W94]

*Type species of generic name indicated


[A27] Allan, R. S. 1927. The geology and palaeontology of the Lower Waihao Basin, south Canterbury, New Zealand. Transactions and Proceedings of the New Zealand Institute 57: 265–309.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[C80] Cernohorsky, W. O. 1980. The taxonomy of some Indo-Pacific Mollusca. Part 8. Records of the Auckland Institute and Museum 17: 135–152.

[CC95] Coovert, G. A., & H. K. Coovert. 1995. Revision of the supraspecific classification of marginelliform gastropods. Nautilus 109 (2–3): 43–110.

[H04] Haeckel, E. 1899–1904. Kunstformen der Natur. Bibliographisches Institut: Leipzig und Wien.

[PB29] Powell, A. W. B., & J. A. Bartrum. 1929. The Tertiary (Waitematan) molluscan fauna of Oneroa, Waiheke Island. Transactions and Proceedings of the New Zealand Institute 60: 395–447.

[PK11] Puillandre, N., Y. I. Kantor, A. Sysoev, A. Couloux, C. Meyer, T. Rawlings, J. A. Todd & P. Bouchet. 2011. The dragon tamed? A molecular phylogeny of the Conoidea (Gastropoda). Journal of Molluscan Studies 77: 259–272.

[R42] Reeve, L. 1842. Descriptions of two new species of Oniscia, a genus of pectinibranchiate mollusks. Proceedings of the Zoological Society of London 10: 91.

[TTE93] Tracey, S., J. A. Todd & D. H. Erwin. 1993. Mollusca: Gastropoda. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 131–167. Chapman & Hall: London.

[W94] Wilson, B. 1994. Australian Marine Shells vol. 2. Prosobranch Gastropods. Part Two (Neogastropods). Odyssey Publishing: Kallaroo (Western Australia).

[WG71] Wilson, B. R., & K. Gillett. 1971. Australian Shells: illustrating and describing 600 species of marine gastropods found in Australian waters. A. H. & A. W. Reed: Sydney.

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