Pleuropoma extincta, copyright Bronwen Scott.

Belongs within: Helicinoidea.

The Helicininae are a group of operculate terrestrial snails in which the radula possesses five lateral teeth, including a comb-lateral and an accessory plate (Knight et al. 1960).

|--Bourciera Pfeiffer 1852BR05 [incl. Pseudhelicina Sykes 1907KC60; Bourcierinae]
| `--*B. helicinaeformis (Pfeiffer 1852) [=Cyclostoma helicinaeforme]BR17
|--Ceratopoma Moellendorff 1893KC60, S88 [incl. Diversicolor Wagner 1905KC60, Negopenia Iredale 1941 (n. n.)KC60]
| `--*C. caroli (Kobelt 1886) [=Helicina caroli]KC60
|--Hemipoma Wagner 1905KC60
| `--*H. hakodadiensis (Hartmann 1890) [=Helicina hakodadiensis]KC60
|--Orobophana Wagner 1905KC60
| `--*O. uberta (Gould 1847) [=Helicina uberta]KC60
|--Sturanya Wagner 1905 [incl. Sturanyella Pilsbry & Cooke 1934, Sturyanella]KC60
| `--*S. plicatilis (Mousson 1865) [=Helicina plicatilis]KC60
|--Palaeohelicina Wagner 1905 (Palaeohelicina)KC60
| |--*P. (P.) fischeriana (Montrouzier 1863) [=Helicina fischeriana]KC60
| `--P. (P.) mayri Clench 1958BC01
|--Olygyra Say 1818BR05 [=OligyraBR05; incl. Subglobulosa Wagner 1905KC60, Succincta Wagner 1905KC60; Oligyradae, Olygyridae]
| |--*O. orbiculata Say 1818 [=Helicina (*Olygyra) orbiculata]KC60
| |--O. oweniana (Pfeiffer 1849) [=Helicina (Oligyra) oweniana; incl. H. coccinostoma Morelet 1849]J68
| `--O. sprucei [=Helicina (Oligyra) sprucei]A66
|--Sulfurina Möllendorff 1893 [incl. Hypostrongyla Tomlin 1930, Pestomena Iredale 1941 (n. n.)]KC60
| |--*S. (Sulfurina) citrina (Grateloup 1840) [=Helicina citrina]KC60
| `--S. (Kosmetopoma Wagner 1905)KC60
| `--S. (*K.) amaliae (Kobelt 1886) [=Helicina amaliae]KC60
|--Schasicheila Shuttleworth 1852 [=Atoyac Baker 1928, Schasichila Fischer 1885]KC60
| |--*S. (Schasicheila) alata (Pfeiffer 1848) [=Helicina alata, *Atoyac alata]KC60
| |--S. (Misantla Baker 1928)KC60
| | `--S. (*M.) misantlensis Fischer & crosse 1893KC60
| `--S. (Necaxa Baker 1928)KC60
| `--S. (*N.) minuscula (Pfeiffer 1859) [=Helicina minuscula]KC60
|--Alcadia Gray 1840 (see below for synonymy)KC60
| | i. s.: A. delmonteana Pilsbry 1932BC01
| | A. fallaxJ65
| | A. pompholyx (Pilsbry 1933) [=Helicina pompholyx]BC01
| |--*A. (Alcadia) major (Gray 1824) [=Helicina major]KC60
| |--A. (Analcadia Wagner 1907)KC60
| | `--A. (*A.) dysoni (Pfeiffer 1849) [=Helicina dysoni]KC60
| |--A. (Emoda Adams & Adams 1856) [incl. Glyptemoda Clench & Aguayo 1950)KC60
| | `--A. (*E.) silacea (Morelet 1849) [=Helicina silacea]KC60
| |--A. (Idesa Adams & Adams 1856) (see below for synonymy)KC60
| | `--A. (*I.) rotunda (d’Orbigny in Sagra 1842) [=Helicina rotunda]KC60
| |--A. (Penisoltia Baker 1954)KC60
| | `--A. (*P.) hispida (Pfeiffer 1839) [=Helicina hispida]KC60
| `--A. (Striatemoda Baker 1940) [incl. Diaphana Guppy 1895 non Brown 1827]KC60
| `--A. (S.) subfusca (Menke 1828) (see below for synonymy)KC60
|--Pleuropoma Moellendorff 1893KC60, KKW12 (see below for synonymy)
| |--*P. dichroa (Möllendorff 1890) [=Helicina dichroa]SS10
| |--P. brecanrio Stanisic in Stanisic, Shea et al. 2010SS10
| |--P. draytonensis (Pfeiffer 1857) [incl. Helicina diversicolor Cox 1866]SS10
| |--P. extincta (Odhner 1917)SS10
| |--P. gladstonensis (Cox 1864)SS10
| |--P. gouldiana (Forbes 1851) (see below for synonymy)SS10
| |--P. jana (Cox 1872)SS10
| |--P. louisiadensis (Forbes 1851)S88
| |--P. macleayi (Brazier 1876)SS10
| |--P. queenslandica Stanisic in Stanisic, Shea et al. 2010SS10
| |--P. reflexilabiata Stanisic in Stanisic, Shea et al. 2010SS10
| |--P. richardson Stanisic in Stanisic, Shea et al. 2010SS10
| |--P. solomonensis (Smith 1885)S88
| |--P. spatei Stanisic in Stanisic, Shea et al. 2010SS10
| |--P. talusata Stanisic in Stanisic, Shea et al. 2010SS10
| |--P. verecunda (Gould 1859) [=Helicina verecunda, P. (*Aphanoconia) verecunda, *Sphaeroconia verecunda]KC60
| `--P. walkeri (Smith 1894)KKW12 (see below for synonymy)
`--Helicina Lamarck 1799BR05 (see below for synonymy)
| i. s.: H. acutissimaS42
| H. chionea Pilsbry 1949BC01
| H. clappiBP90
| H. guadeloupensisS42
| H. hollandi Adams 1849BC01
| H. laciniosa Mighels 1845 [=Pleuropoma laciniosa; incl. Orobophana berniceia]J49
| H. lazarusS42
| H. lindeniC64
| H. malleataG79b
| H. maximaS42
| H. merdigeraC64
| H. minutaS42
| H. parvaS42
| H. politaS42
| H. rotelloidea Mighels 1845 [incl. Pleuropoma hawaiiensis]J49
| H. rugosaG79b
| H. salleana Pfeiffer 1852WSS20
| H. similisS42
| H. striatulaS42
| H. substriataG79a
| H. trochiformisS42
| H. turbinataC64
|--*H. (Helicina) neritella Lamarck 1801 [=Helix neritella, *Euneritella neritella, *Pitonnellus neritella]KC60
|--H. (Oxyrhombus Fischer & Cross 1893)KC60 (see below for synonymy)
| |--H. (*O.) amoena Pfeiffer 1849KC60, J68 [=*Ampullina amoenaCJ66]
| |--H. concentricaCJ66 [=Ampullina concentricaCJ66, H. (Pachystoma) concentricaA66]
| `--H. moquiniana [=Ampullina moquiniana]CJ66
|--H. (Pseudoligyra Baker 1954)KC60
| `--H. (*P.) tenuis Pfeiffer 1849KC60
`--H. (Tristramia Crosse 1863) [incl. Caloplisma Fischer & Crosse 1893, Retorquata Wagner 1905]KC60
|--H. (*T.) salvini Tristram 1861KC60
`--H. (T.) rostrata Morelet 1851J68

Alcadia Gray 1840 [incl. Eualcadia Wagner 1907, Incrustata Wagner 1907, Intusplicata Wagner 1907, Isoltia Guppy 1859, Palliata Wagner 1907, Sericea Wagner 1907]KC60

Alcadia (Idesa Adams & Adams 1856) [incl. Hjalmarsona Baker 1940, Leialcadia Wagner 1907, Mammilla Wagner 1907 non Schumacher 1817, Schrammia Guppy 1895, Weinlandella Baker 1954]KC60

Alcadia (Striatemoda) subfusca (Menke 1828) [=Helicina subfusca; incl. H. striata Lamarck 1822 non Defrance 1821, Alcadia (*Striatemoda) striata]KC60

Helicina Lamarck 1799BR05 [=Euneritella Wagner 1905KC60, Pitonnellus deMontfort 1810KC60, Pitonnillus deMontfort 1810KC60; incl. Ampliata Wagner 1907KC60, Colyma Rafinesque 1815KC60, Festiva Wagner 1910KC60, Pachytoma Swainson 1840KC60]

Helicina (Oxyrhombus Fischer & Cross 1893)KC60 [=Ampullina Guppy 1895 nec Bowdich 1822 nec deBlainville 1824 nec Quél. 1875 (ICBN)CJ66; incl. Angulata Wagner 1905KC60, Tamsiana Wagner 1905KC60]

Pleuropoma Moellendorff 1893KC60, KKW12 [incl. Albocincta Wagner 1909SS10, Aphanoconia Wagner 1905SS10, Dichroa Wagner 1905SS10, Pachystoma Wagner 1905 non Guilding 1828KC60, Reticulata Wagner 1905SS10, Sculpta Wagner 1905SS10, Sphaeroconia Wagner 1909SS10]

Pleuropoma gouldiana (Forbes 1851) [incl. Helicina fumigata Sowerby 1866, H. lizardensis Cox 1868, H. reticulata Pfeiffer 1863, Aphanoconia (Sphaeroconia) subreticulata Wagner 1909]SS10

Pleuropoma walkeri (Smith 1894)KKW12 [=Helicina walkeriKKW12, Orobophana walkeriS88, Aphanoconia (Sphaeroconia) baudinensis Wagner 1909KKW12]

*Type species of generic name indicated


[A66] Adams, H. 1866. List of land and freshwater shells collected by Mr. E. Bartlett on the Upper Amazons, and on the River Ucayali, eastern Peru, with descriptions of new species. Proceedings of the Zoological Society of London 1866: 440–445.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[BP90] Burch, J. B., & T. A. Pearce. 1990. Terrestrial Gastropoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 201–309. John Wiley & Sones: New York.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[CJ66] Clench, W. J., & M. K. Jacobson. 1966. On Ampullina Guppy. Nautilus 80 (2): 71.

[G79a] Gibbons, J. S. 1879a. Notes on the habits and distribution, &c., of certain W. Indian Pulmonifera. Journal of Conchology 2: 129–134.

[G79b] Gibbons, J. S. 1879b. Land shells collected at Puerto Plata, San Domingo. Journal of Conchology 2: 209.

[J65] Jacobson, M. K. 1965. On some land shells of Eleuthera, Bahamas. Nautilus 78 (4): 120–125.

[J68] Jacobson, M. K. 1968. On a collection of terrestrial mollusks from Nicaragua. Nautilus 81 (4): 114–119.

[J49] Johnson, R. I. 1949. Jesse Wedgwood Mighels with a bibliography and a catalogue of his species. Occasional Papers on Mollusks 1 (14): 213–231.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[KKW12] Köhler, F., V. Kessner & C. Whisson. 2012. New records of non-marine, non-camaenid gastropods (Mollusca: Gastropoda) from islands off the Kimberley coast, Western Australia. Records of the Western Australian Museum 27 (1): 21–39.

[S88] Solem, A. 1988. Non-camaenid land snails of the Kimberley and Northern Territory, Australia. I. Systematics, affinities and ranges. Invertebrate Taxonomy 4: 455–604.

[S42] Sowerby, G. B., Jr. 1842. Descriptions of some new species of Helicinae, in the collection of H. Cuming, Esq. Proceedings of the Zoological Society of London 10: 6–8.

[SS10] Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A. field guide to eastern Australian species. Bioculture Press: Mauritius.

[WSS20] Watters, G. T., M. L. Smith & D. J. Sneddon. 2020. The subfamily Abbottellinae (Gastropoda: Annulariidae): origins, associations, and a review of the Hispaniolan taxa. Nautilus 134 (1): 1–34.

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