Helix

Edible snail Helix pomatia, copyright Björn S.

Belongs within: Helicinae.

Helix is a western Palaearctic genus including the edible snail H. pomatia and related species.

<==Helix Linnaeus 1758 [=Pentataenia Schmidt 1855; Pentataeniae, Pentataeniidae, Pentataeniinae]BR05
    |--*H. (Helix) pomatia Linnaeus 1758BR17 (see below for synonymy)
    |--H. (Aglaia) infumataC64
    |--H. (Camoena) piliodon Benson 1860TC89
    |--H. (Carocolla)S41d
    |    |--‘Carocolla’ bidentataG40
    |    |--H. (C.) dealbata Broderip 1841B41c
    |    |--H. (C.) dryope Broderip 1841B41b
    |    |--H. (C.) listeri Broderip 1841 [incl. Carocolla gallina]B41b
    |    |--H. (C.) panayensis Brod. in Pfeiffer 1842P42a
    |    |--H. (C.) papyracea Broderip 1841B41b
    |    |--H. (C.) parmula Broderip 1841B41b
    |    |--H. (C.) puella Broderip 1841B41c
    |    |--H. (C.) reginae Broderip 1841B41b
    |    |--H. (C.) rota Broderip 1841B41c
    |    |--H. (C.) semigranosa Sowerby 1841S41d
    |    |--H. (C.) siquijorensis Broderip 1841B41b
    |    |--H. (C.) thersites Broderip 1841B41b
    |    |--H. (C.) virgo Broderip 1841B41c
    |    `--H. (C.) zebuensis Broderip 1841B41c
    |--H. (Genesella) hariolaTC89
    |--H. (Heliomanes)G40
    |    |--H. (H.) caperataG40 [=Teba caperataG40; incl. H. crenulataG40, H. intersectaG40; incl. H. caperata var. ornataA79]
    |    `--H. (H.) ericetorum [=Zonites ericetorum; incl. H. erica, H. obliterata]G40
    |--H. (Hygromanes)G40
    |    |--H. (H.) cantiana [=Teba cantiana]G40
    |    |--H. (H.) carthusiana (see below for synonymy)G40
    |    |--H. (H.) concinna Sowerby 1841G40, S41c
    |    |--H. (H.) depilataG40
    |    |--H. (H.) fusca [incl. H. (Zenobia) corrugata Gray 1821, H. subrufescens]G40
    |    |--H. (H.) lamellata [incl. H. scarburgensis]G40
    |    |--H. (H.) limbata [incl. H. circinata]G40
    |    |--H. (H.) revelataG40
    |    |--H. (H.) rufescensG40 (see below for synonymy)
    |    `--H. (H.) sericeaG40
    |--H. (Plectotropis)TC89
    |    |--H. (P.) fulvicans Adams 1866A66
    |    |--H. (P.) huttoni [incl. H. orbicula]TC89
    |    |--H. (P.) tapeina Benson 1836TC89
    |    `--H. (P.) theobaldiana Tapparone Canefri 1889 [=H. rotatoria Theobal 1859 (preoc.)]TC89
    |--H. (Tapada Gray 1840)G40
    |    `--H. (*T.) aperta [incl. H. naticoides, H. neritoides]G40
    `--H. (Trachia)TC89
         |--H. (T.) delibrata Benson 1836 [incl. H. procumbens Gould 1844]TC89
         `--H. (T.) gabata [incl. H. merguiensis Philippi 1846]TC89
Helix incertae sedis:
  H. acuminata Sowerby 1841S41e
  H. albolabris [incl. H. albolabris var. dentata]W79
  H. ambigua Linnaeus 1758L58
  H. arboreaC64
  H. asteriscusW79
  H. auriculataB41b
  H. balthica Linnaeus 1758L58
  H. barrenensis Petterd 1879P79b
  H. bassiP79b
  H. beckiana Pfeiffer 1842P42a
  H. bisulcataL79
  H. bolivariE99
  H. brevidens Sowerby 1841S41d
  H. brevipilaC66
  H. breweriC64
  H. brunnea Sowerby 1841S41e
  H. bulbulusC84
  H. bulla Pfeiffer 1842P42b
  H. butleri Pfeiffer 1842P42a
  H. caespitumG40
  H. caesusP79b
  H. cailliaudiS41a
  H. candidulaG40
  H. capitiumTC89
  H. carbonaria Sow. in Pfeiffer 1842P42a
  H. carpenterianaC64
  H. cazenarettiSm79
  H. chersinaC64
  H. chlorochroa Sowerby 1841S41a
  H. chrysocheilus Sowerby 1841S41a
  H. cincinniformis Sowerby 1841S41b
  H. clausaW79
  H. coccomelosS41a
  H. columbaria Sowerby 1841S41c
  H. conscendens Cox 1866C66
  H. conspurcataG40
  H. cornuarietis Linnaeus 1758L58
  H. cornumilitare Linnaeus 1758L58
  H. corticicola Cox 1866C66
  H. crebristriataC64
  H. cretata Broderip 1841B41a
  H. cromyodes Pfeiffer 1842P42b
  H. cryptica Broderip 1841B41a
  H. cubensisG79a
  H. cultellataC64
  H. cumingii Pfeiffer 1842P42a
  H. curacoaeP79b
  H. curta Sowerby 1841S41c
  H. dandenongensis Petterd 1879P79c
  H. diversicolor Cox 1866C66
  H. elevataW79
  H. engaddensisH84
  H. euryomphalaC64
  H. exceptiunculaC84
  H. eximiaC64
  H. exoletaW79
  H. fallaxW79
  H. fasciolataSc79
  H. fernshawensis Petterd 1879P79c
  H. figulinaC63
  H. fulgens Sowerby 1841S41a
  H. fulvida Pfeiffer 1842P42a
  H. furneauxensis Petterd 1879P79b
  H. gabbiiC64
  H. glauca Linnaeus 1758L58
  H. gothica Linnaeus 1758L58
  H. grisea Linnaeus 1758L58
  H. gualtierana Linnaeus 1758L58
  H. gummata Sowerby 1841S41d
  H. halliP79b
  H. hamiltoniP79b
  H. harfordiiB41a
  H. harpa Say 1844 [incl. Pupa costulata Mighels 1844]J49
  H. henryana Petterd 1879P79b
  H. hillebrandi Newcomb 1864C64
  H. hirsutaW79
  H. holosericeaG40
  H. hystrix Mighels in Pfeiffer 1846J49
  H. incarnataG40
  H. incomptaS41e
  H. indistinctaG79b
  H. inflectaW79
  H. janus [=Helicella janus; incl. Helix mackensiana Saul 1841]P42a
  H. jugosa Mighels 1845J49
  H. jungermanniae Petterd 1879P79b
  H. kershawi Petterd 1879P79b
  H. lallianaC64
  H. lanculaSm79
  H. languida Pfeiffer 1842P42b
  H. laomontanaTC89
  H. latitans Broderip 1841B41a
  H. laxataSm79
  H. legrandiP79b
  H. leidyi Hall & Meek 1856W77
  H. leucas Linnaeus 1758L58
  H. leucophaea Sowerby 1841S41b
  H. limulaP79c
  H. lottah Petterd 1879P79b
  H. loxodonG79b
  H. lucorum Linnaeus 1758L58 [incl. H. castaneaG40, H. mutataG40]
  H. lutaria Linnaeus 1758L58
  H. luzonica Sow. in Pfeiffer 1842P42a
  H. macgillivrayiM86
  H. maculosa [incl. H. irregularis]G40
  H. mathinnae Petterd 1879P79b
  H. macdonaldiT79
  H. mammillaP42b
  H. matruelis Sowerby 1841S41d
  H. meretrix Sowerby 1841S41c
  H. metaformisS41a
  H. mimosa Petterd 1879P79b
  H. mindanaensis Sow. in Pfeiffer 1842P42a
  H. modesta Sowerby 1841S41e
  H. monochroa Sowerby 1841S41a
  H. monodon [incl. H. monodon var. fraterna, H. monodon var. leaii]W79
  H. moricandi Sow. in Pfeiffer 1842P42a
  H. multilineata [incl. H. multilineata var. unicolor]W79
  H. murrayanaT79
  H. nelsonensisP79b
  H. neritoidea Linnaeus 1758L58
  H. newberryanaC64
  H. norrisii Sow. in Pfeiffer 1842P42a
  H. oblonga Sowerby 1841S41e
  H. occidentalisSc79
  H. oculuscapri Linnaeus 1758L58
  H. olivieriG40
  H. oomorphaS41e
  H. otahaitanaJ42
  H. palliata [incl. H. palliata var. alba non H. alternata var. alba]W79
  H. pan Broderip 1841B41a
  H. parvissimaP79b
  H. peguensisTC89
  H. peregrinaC64
  H. perspicua Linnaeus 1758L58
  H. petterdiana Taylor 1879 [incl. H. petterdiana var. albida Taylor 1879]T79
  H. phloiodes Pfeiffer 1842P42b
  H. pictilisP79a
  H. plebeiaG40
  H. polygyrataC64
  H. porphyria Pfeiffer 1842P42a
  H. pulcherrimaP42a
  H. pyramidalis Sowerby 1841S41e
  H. pytyonesicaC64
  H. radiansP79c
  H. ravidaC42
  H. reevii Broderip 1841B41b
  H. roblini Petterd 1879P79b
  H. roissyanaS41d
  H. rosacea Petterd 1879P79b
  H. rothiC64
  H. rufocinctaC64
  H. runssorinaPB27
  H. sagittifera Pfeiffer 1842P42a
  H. sagraianaC64
  H. samarensis Pfeiffer 1842P42a
  H. sayiiW79
  H. scarabaeus Linnaeus 1758L58
  H. scrobiculata Pfeiffer 1842P42a
  H. setiger Sowerby 1841S41d
  H. setipilaTC89
  H. solitariaW79
  H. sphaerica Sowerby 1841S41d
  H. sphaerion Sowerby 1841S41a
  H. stephensiP79b
  H. striatellaW79
  H. striatula Linnaeus 1758L58
  H. subangulataL79
  H. subaquilaG79a
  H. subplicatusCC06
  H. subrutila Mighels 1845J49
  H. sylvatica [incl. H. austriaca, H. vindebonensis]G40
  H. tamarensisT79
  H. tasmaniaeT79
  H. tephrodes Pfeiffer 1842P42b
  H. terrestrisSc79
  H. thyroidesW79
  H. tiara Mighels 1845J49
  H. trucanini Petterd 1879P79b
  H. tryoniC64
  H. unica Pfeiffer 1842P42b
  H. urarensisC66
  H. valtoni Reeve 1842R42
  H. varians [incl. H. submeris Mighels 1844]J49
  H. versicolorP42a
  H. vilanovaiE99
  H. vitrinaeformisP79c
  H. wellingtonensisP79a
  H. whitneyiC64
  H. wynyardensis Petterd 1879P79b
  H. xysteraSm79
  H. zeus Jonas 1842J42
  H. zonalisP42a
  H. zonifera Sow. in Pfeiffer 1842P42a

Nomen nudum: Helix sandiegoensis Lea in Gould 1855C64

*Helix (Helix) pomatia Linnaeus 1758BR17 [=*Pentataenia pomatiaBR17, Cochlea pomatiaG40; incl. Pomatia antiquorumG40, C. edulisG40, Helix pomanaG40, H. scalarisG40]

Helix (Hygromanes) carthusiana [incl. H. (Zenobia) bimarginata Gray 1821, H. carthusianella, Teba carthusianella, H. gibbsii, H. gypsii (l. c.), H. rufilabris]G40

Helix (Hygromanes) rufescensG40 [=Teba rufescensG40; incl. H. rufescens var. albida non H. petterdiana var. albida Taylor 1879H79, H. glabellaG40]

*Type species of generic name indicated

References

[A66] Adams, H. 1866. Descriptions of fifteen new species of land and freshwater shells from Formosa, collected by Robert Swinhoe, Esq., consul at Taiwan in that island. Proceedings of the Zoological Society of London 1866: 316-319.

[A79] Ashford, C. 1879. Land and freshwater shells observed in the neighborhood of Redcar. Journal of Conchology 2: 236–241.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BR17] Bouchet, P., J.-P. Rocroi, B. Hausdorf, A. Kaim, Y. Kano, A. Nützel, P. Parkhaev, M. Schrödl & E. E. Strong. 2017. Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia 61 (1–2): 1–526.

[B41a] Broderip, W. J. 1841a. Descriptions of shells collected by H. Cuming, Esq., in the Philippine Islands. Proceedings of the Zoological Society of London 9: 22–23.

[B41b] Broderip, W. J. 1841b. Descriptions of shells collected by H. Cuming, Esq., in the Philippine Islands (continued). Proceedings of the Zoological Society of London 9: 34–39.

[B41c] Broderip, W. J. 1841c. Descriptions of shells collected by H. Cuming, Esq., in the Philippine Islands (continued). Proceedings of the Zoological Society of London 9: 44–46.

[C84] Cain, A. J. 1984. Heterosematism in snails. Malacologia 25 (1): 161–172.

[CC06] Cameron, R. A. D., L. M. Cook, G. A. Goodfriend & M. B. Seddon. 2006. Fossil land snails of Porto Santo, Madeiran archipelago: change and stasis in Pleistocene to Recent times. Malacologia 49 (1): 25–59.

[C42] Cantor, D. 1842. Letter containing a list of a collection of shells from Chusan, presented to the society. Proceedings of the Zoological Society of London 10: 124.

[C64] Carpenter, P. P. 1864. Supplementary report on the present state of our knowledge with regard to the Mollusca of the west coast of North America. Report of the British Association for the Advancement of Science 33: 517–686.

[C66] Cox, J. C. 1866. Characters of six new Australian land-shells. Proceedings of the Zoological Society of London 1866: 373–375.

[C63] Curtin, T. J. 1963. Snails from Turkey, Crete and Greece. Nautilus 76 (4): 132–135.

[E99] Esu, D. 1999. Contribution to the knowledge of Neogene climate changes in western and central Europe by means of non-marine molluscs. In: J. Agustí, L. Rook & P. Andrews (eds) The Evolution of Neogene Terrestrial Ecosystems in Europe pp. 328–354. Cambridge University Press.

[G79a] Gibbons, J. S. 1879a. Notes on the habits and distribution, &c., of certain W. Indian Pulmonifera. Journal of Conchology 2: 129–134.

[G79b] Gibbons, J. S. 1879b. Land shells collected at Puerto Plata, San Domingo. Journal of Conchology 2: 209.

[G40] Gray, J. E. 1840. A Manual of the Land and Fresh-water Shells of the British Islands, with figures of each of the kinds. By William Turton, M.D. A new edition, thoroughly revised and much enlarged. Longman, Orme, Brown, Green, and Longmans: London.

[H84] Heller, J. 1984. Shell colours of desert landsnails. Malacologia 25 (2): 355–359.

[H79] Hey, W. C. 1879. Contributions to a better knowledge of the land and freshwater shells of Yorkshire. Journal of Conchology 2: 310–314.

[J49] Johnson, R. I. 1949. Jesse Wedgwood Mighels with a bibliography and a catalogue of his species. Occasional Papers on Mollusks 1 (14): 213–231.

[J42] Jonas, J. H. 1842. Descriptions of new species of shells belonging to the genera Helix and Bulimus, collected by Mr. Cuming in the Philippine Islands. Proceedings of the Zoological Society of London 10: 188–189.

[L79] Legrand, W. 1879. Notes on a few Tasmanian land and freshwater shells. Journal of Conchology 2: 95–96.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[M86] Masters, G. 1886. Notes and exhibits. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 477.

[P79a] Petterd, W. F. 1879a. List of the freshwater shells of Tasmania. Journal of Conchology 2: 80–88.

[P79b] Petterd, W. F. 1879b. Descriptions of some new Tasmanian Helices. Journal of Conchology 2: 210–217.

[P79c] Petterd, W. F. 1879c. Descriptions of three new Helices from Australia. Journal of Conchology 2: 355–356.

[P42a] Pfeiffer, L. 1842a. Descriptions of new species of shells belonging to the genera Helix and Bulimus, collected by H. Cuming, Esq. in the Philippine Islands. Proceedings of the Zoological Society of London 10: 84–89.

[P42b] Pfeiffer, L. 1842b. Descriptions of new species of shells, collected by Hugh Cuming, Esq. in the Philippine Islands. Proceedings of the Zoological Society of London 10: 150–153.

[PB27] Pilsbry, H. A., & J. Bequaert. 1927. The aquatic mollusks of the Belgian Congo, with a geographical and ecological account of Congo malacology. Bulletin of the American Museum of Natural History 53 (2): 69–602, pls 10–77.

[R42] Reeve, L. 1842. Descriptions of new species of shells, principally from the collection of Hugh Cuming, Esq. Proceedings of the Zoological Society of London 10: 49–50.

[Sc79] Scharff, R. 1879. Land and freshwater shells collected in the alluvium of the Garonne, France. Journal of Conchology 2: 315–316.

[Sm79] Smith, E. A. 1879. Descriptions to two species of Helicidae from Madagascar. Journal of Conchology 2: 337–339.

[S41a] Sowerby, G. B. 1841a. Descriptions of new species of shells, collected by H. Cuming, Esq., in the Philippine Islands. Proceedings of the Zoological Society of London 9: 1–3.

[S41b] Sowerby, G. B. 1841b. On species of Helix collected by H. Cuming, Esq., in the Philippine Islands. Proceedings of the Zoological Society of London 9: 17–18.

[S41c] Sowerby, G. B. 1841c. Descriptions of new species of shells, collected by H. Cuming, Esq., in the Philippine Islands (continued). Proceedings of the Zoological Society of London 9: 19–20.

[S41d] Sowerby, G. B. 1841d. Descriptions of new species of shells, collected by H. Cuming, Esq., in the Philippine Islands (continued). Proceedings of the Zoological Society of London 9: 24–26.

[S41e] Sowerby, G. B. 1841e. Descriptions of new species of shells, collected by H. Cuming, Esq., in the Philippine Islands (continued). Proceedings of the Zoological Society of London 9: 39–40.

[TC89] Tapparone Canefri, T. 1889. Viaggio di Leonardo Fea in Birmania e regioni vicine. XVIII.—Molluschi terrestri e d’acqua dolce. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 295–359.

[T79] Taylor, J. W. 1879. Description of Helix petterdiana, new species. Journal of Conchology 2: 287–288.

[W79] Walker, B. 1879. Catalogue of the shell-bearing Mollusca of Michigan. Journal of Conchology 2: 325–337.

[W77] White, C. A. 1877. Report upon the invertebrate fossils collected in portions of Nevada, Utah, Colorado, New Mexico, and Arizona, by parties of the expeditions of 1871, 1872, 1873, and 1874. U.S. Geographical Surveys West of the One Hundredth Meridian 4 (1): 1–219, pls 1–21.

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