Hemileiidae

Dorsal (left) and ventral views of Hemileius paraguayensis, from Balogh & Balogh (1992b).

Belongs within: Oripodoidea.
Contains: Hemileius (Hemileius), Hemileius (Tuberemaeus).

The Hemileiidae are a group of oripodoid mites with a sacculate octotaxic system and lacking pteromorphs (Balogh & Balogh 1992).

Characters (from Balogh & Balogh 1992a, as Hemileius group of Scheloribatidae): Octotaxic system with sacculi; pteromorphae absent. Usually four pairs of genital setae, exceptionally five; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae.

<==Hemileiidae [Hemileinae]S04
    |--Constrictobates Balogh & Mahunka 1966 [Constrictobatinae]S04
    |    `--*C. lineolatus Balogh & Mahunka 1966S04
    |--Mucrobates Balogh & Mahunka 1979S04
    |    `--*M. fissuratus Balogh & Mahunka 1979S04
    |--Plumobates Balogh & Mahunka 1966S04
    |    `--*P. decoratus Balogh & Mahunka 1966S04
    |--Zeascheloribates Luxton 1982S04
    |    `--*Z. palustris Luxton 1982S04
    |--Vesiculobates Hammer 1979S04
    |    `--*V. silvatica Hammer 1979S04, EA14 [=Siculobata (*Vesiculobates) silvaticaS04]
    |--Baobabula Mahunka 1975S04
    |    |--*B. mussardi Mahunka 1975S04
    |    `--B. impolita (Warburton 1912) [=Notaspis impolita]S04
    |--Monoschelobates Balogh & Mahunka 1969S04
    |    |--*M. parvus Balogh & Mahunka 1969S04
    |    `--M. translamellatus Pérez-Íñigo & Baggio 1991S04
    |--Balazsella Mahunka 1983S04
    |    |--*B. pilososetosa Mahunka 1983Mah83
    |    |--B. ilhabellae Balogh & Palacios-Vargas 1996S04
    |    `--B. mexicana Balogh & Palacios-Vargas 1996S04
    |--Heteroleius Balogh & Mahunka 1966S04
    |    |--*H. longissimus Balogh & Mahunka 1966S04
    |    |--H. navicula (Berlese 1913) [=Oribatula navicula]S04
    |    `--H. planus (Aoki 1984) [=Symphauloppia plana]S04
    |--Nasozetes Sellnick 1930 [incl. Kinabaluella Mahunka 1966]S04
    |    |--*N. sumatrensis Sellnick 1930S04
    |    |--N. choerognathus Willmann 1931S04
    |    |--N. porcella (Mahunka 1996) [=Kinabaluella porcella]S04
    |    `--N. stunkardi Sengbusch 1957S04
    |--Simkinia Krivolutsky 1966S04
    |    |--*S. turanica Krivolutsky 1966 [incl. Hemileius ovalis Kulijev 1968]S04
    |    |--S. elongata Krivolutsky 1969S04
    |    |--S. montana Krivolutsky & Grishina 1970S04
    |    |--S. schachtachtinskoi (Kulijev 1961) [=Oribatula schachtachtinskoi]S04
    |    `--S. tianshanica Krivolutsky 1971S04
    |--Siculobata Grandjean 1953 [incl. Floribates Norton & Kethley 1989]S04
    |    |--*S. sicula (Berlese 1892)S04 [=Oppia tibialis siculaS04, Notaspis siculaM98, Oribatula (Hemileius) siculaB16]
    |    |    |--S. s. siculaS04
    |    |    `--S. s. platensis (Berlese 1916)S04 [=Oribatula (Hemileius) sicula var. platensisB16]
    |    |--S. florens (Berlese 1908) [=Oribatula florens]S04
    |    |--S. malabarica Ramani & Haq 1998S04
    |    `--S. talarica Fredes & Martinez 2013EA14
    |--Exoribatula Jacot 1936S04
    |    |  i. s.: E. flagelligera (Balogh 1958)B65
    |    |--E. (Exoribatula) [incl. Nesoribatula Aoki 1964]S04
    |    |    |--*E. (E.) biundata Jacot 1936S04 [=E. biundatusAO74]
    |    |    |--E. (E.) ensifer (Mahunka 1983) [=Nesoribatula ensifer]S04
    |    |    |--E. (E.) pacifica (Aoki 1964)S04 [=*Nesoribatula pacificaA64]
    |    |    `--E. (E.) rotunda (Mahunka 1988) [=Nesoribatula rotunda]S04
    |    `--E. (Multoribates Hammer 1961)S04
    |         |--E. (*M.) chavinensis (Hammer 1961) [=*Multoribates chavinensis]S04
    |         |--E. (M.) longior (Hammer 1958) [=Scheloribates longior]S04
    |         |--‘Multoribates’ minorMar83
    |         |--E. (M.) multisetosus (Balogh 1970) [=Tuberemaeus multisetosus]S04
    |         `--E. (M.) parvialatus (Hammer 1958) [=Scheloribates parvialatus]S04
    `--Hemileius Berlese 1916S04
         |--H. (Hemileius)S04
         |--H. (Tuberemaeus)S04
         |--H. (Turcibates Ayyildiz & Luxton 1989) [incl. Tenuileius Lee 1989]S04
         |    |--H. (*T.) parvus (Ayyildiz & Luxton 1989) (see below for synonymy)S04
         |    |--H. (T.) lanceolatus Bayartogtokh & Aoki 1997S04
         |    |--H. (T.) minimus Lee 1989S04 [=H. (*Tenuileius) minimusBB92a]
         |    |--H. (T.) paratenuis Lee 1989 [=H. (Tenuileius) paratenuis]S04
         |    `--‘Turcibates’ porosus Ayyildiz & Luxton 1989BB92b
         `--H. (Urubambates Hammer 1961)S04
              |--H. (*U.) punctatus (Hammer 1961) [=*Urubambates punctatus]S04
              |--H. (U.) calcaratus (Mahunka 1984) [=Urubambates calcaratus]S04
              |--H. (U.) flagellatus (Mahunka 1985) [=Urubambates flagellatus]S04
              |--H. (U.) paraguayensis (Balogh & Mahunka 1981) [=Urubambates paraguayensis]S04
              |--H. (U.) perlongus (Vasiliu & Calugar 1976) [=Urubambates perlongus]S04
              `--H. (U.) romanicus (Vasiliu & Calugar 1981) [=Urubambates romanicus]S04

Hemileius (*Turcibates) parvus (Ayyildiz & Luxton 1989) [=*Turcibates parvus; incl. Dometorina rossica Bashkirova 1958 (n. d.)]S04

*Type species of generic name indicated

References

[A64] Aoki, J. 1964. Some oribatid mites (Acarina) from Laysan Island. Pacific Insects 6 (4): 649–664.

[AO74] Aoki, J., & N. Ohkubo. 1974. A proposal of new classification of the family Oripodidae (s. lat.), with description of new species. Bulletin of the National Science Museum, Tokyo 17 (2): 117–147.

[B65] Balogh, J. 1965. A synopsis of the world oribatid (Acari) genera. Acta Zoologica Academiae Scientiarum Hungaricae 11 (1–2): 5–99.

[BB92a] Balogh, J., & P. Balogh. 1992a. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[BB92b] Balogh, J., & P. Balogh. 1992b. The Oribatid Mites Genera of the World vol. 2. Plates. Hungarian Natural History Museum: Budapest.

[B16] Berlese, A. 1916. Centuria terza di Acari nuovi. Redia 12: 289–338.

[EA14] Ermilov, S. G., & A. E. Anichkin. 2014. A new species of Scheloribates (Scheloribates) from Vietnam, with notes on taxonomic status of some taxa in Scheloribatidae (Acari, Oribatida). International Journal of Acarology 40 (1): 109–116.

[Mah83] Mahunka, S. 1983. Data to the knowledge of the oribatid fauna of Surinam and Brasil (Acari). Folia Entomologica Hungarica 44 (2): 205–227.

[Mar83] Martin, N. A. 1983. Miscellaneous observations on a pasture fauna: an annotated species list. DSIR Entomology Division Report 3: 1–98.

[M98] Michael, A. D. 1898. Oribatidae. In: H. Lohmann (ed.) Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen vol. 3. Acarina pp. 1–93. R. Friedländer und Sohn: Berlin.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

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