Hemileius (Tuberemaeus)

Dorsal (left) and ventral views of Hemileius areolatus, from Balogh & Balogh (1992 vol. 2).

Belongs within: Hemileiidae.

The subgenus Tuberemaeus of the genus Hemileius is a pantropical group of ornamented oribatid mites.

Characters (from Balogh & Balogh 1992): Prodorsum with lamellae present, translamella absent. Dorsosejugal suture arched. Notogaster foveolate, punctate or reticulate; with ten or fourteen pairs of distinct setae; four pairs of sacculi; pteromorphae absent. Four pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Legs tridactyle.

<==Hemileius (Tuberemaeus Sellnick 1930) [incl. Anisochthodes Newell 1957, Setulobates Mahunka 1984; Tuberemaeinae]S04
    |--H. (*T.) singularis (Sellnick 1930) [=*Tuberemaeus singularis]S04
    |--H. (T.) areolatus (Balogh & Mahunka 1967) [=Tuberemaeus areolatus]S04
    |--H. (T.) asiaticus (Yamamoto & Aoki 2000) [=Tuberemaeus asiaticus]S04
    |--H. (T.) bellissimus (Hammer 1971) [=Tuberemaeus bellissimus]S04
    |--H. (T.) deletus (Hammer 1979) [=Tuberemaeus deletus]S04
    |--H. (T.) dispar (Balogh & Mahunka 1974) [=Tuberemaeus dispar]S04
    |--H. (T.) fissuratus (Balogh 1970) [=Tuberemaeus fissuratus]S04
    |--H. (T.) formosanus (Tseng 1984) [=Tuberemaeus formosanus]S04
    |--H. (T.) foveolatus (Balogh 1958) [=Liebstadia foveolata]S04
    |    |--H. f. foveolatusS04
    |    `--H. f. tridactylus (Balogh 1959) [=Liebstadia foveolata tridactyla]S04
    |--H. (T.) glaber (Balogh 1970) [=Tuberemaeus glaber]S04
    |--H. (T.) heterotrichus (Mahunka 1984) [=Setulobates heterotrichus]S04
    |--H. (T.) indentatus (Hammer 1973) [=Tuberemaeus indentatus]S04
    |--H. (T.) inornatus (Pérez-Íñigo 1982) [=Tuberemaeus inornatus]S04
    |--H. (T.) lagunensis (Corpuz-Raros 1979) [=Tuberemaeus lagunensis]S04
    |--H. (T.) lineatus (Balogh 1970) [=Tuberemaeus lineatus]S04
    |--H. (T.) longisetosus (Balogh 1970) [=Tuberemaeus longisetosus]S04
    |--H. (T.) luzonensis (Corpuz-Raros 1979) [=Tuberemaeus luzonensis]S04
    |--H. (T.) nagaii (Mahunka 1988) [=Tuberemaeus nagaii]S04
    |--H. (T.) neonominatus Subías 2004 [=Tuberemaeus punctatus Balogh 1970 non Urubambates punctatus Hammer 1961]S04
    |--H. (T.) papillifer (Newell 1957) [=Anisochthodes papillifer]S04
    |--H. (T.) perforatoides (Hammer 1979) [=Tuberemaeus perforatoides]S04
    |--H. (T.) perforatus (Willmann 1931) [=Liebstadia perforata]S04
    |--H. (T.) propinquus (Balogh & Mahunka 1974) [=Tuberemaeus propinquus]S04
    |--H. (T.) pseudoareolatus (Mahunka 1977) [=Tuberemaeus pseudoareolatus]S04
    |--‘Tuberemaeus’ reticulatus (Oudemans 1916)C-R79
    |--H. (T.) scrobinus (Berlese 1916) [=Oribatula (Hemileius) scrobina]S04
    |--H. (T.) sculpturatus (Mahunka 1987) [=Tuberemaeus sculpturatus]S04
    |--H. (T.) similis (Balogh 1970) non Notaspis similis Michael 1888 (not preoc. if in dif. gen) [=Tuberemaeus similis]S04
    |--H. (T.) subareolatus (Mahunka 1977) [=Tuberemaeus subareolatus]S04
    |--H. (T.) thienemanni (Willmann 1931) [=Liebstadia thienemanni]S04
    |--H. (T.) tuberculatus (Balogh 1970) [=Tuberemaeus tuberculatus]S04
    `--H. (T.) vermiculatus (Balogh 1970) [=Tuberemaeus vermiculatus]S04

*Type species of generic name indicated


Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[C-R79] Corpuz-Raros, L. A. 1979. Philippine Oribatei (Acarina). I. Preliminary list of species and descriptions of forty new species. Philippine Agriculturist 62 (1): 1–82.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

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