Paracalais suboculatus, copyright Bill & Mark Bell.

Belongs within: Agrypninae.

The Hemirhipini are a group of click beetles with scale-like setae and the medial margin of the hypomeron evenly margined (Johnson 2002).

<==Hemirhipini [Chalcolepidiini]KB11
    |--Calais speciosus (Linnaeus 1767)B14
    |--Cryptaulus larvatus (Candèze 1874)KB11, PS07 [=Alaus larvatusPS07]
    |--Pherhimius Fleutiaux 1942J02
    |--Tetrigus Candèze 1857PS07, C96
    |    |--*T. parallelus Candèze 1857C96
    |    |--T. australicus Blackburn 1896C96
    |    |--T. lewisi Candèze 1873PS07
    |    `--T. murrayi Waterhouse 1900JG19
    |--Chalcolepidius Eschscholtz 1829KB11, B14
    |    |--C. limbatus Eschscholtz 1829B14
    |    |--C. obscurus Cast. 1836 [incl. C. defloratus, C. lherminieri]FS90
    |    |--C. porcatusI92
    |    `--C. tartarusJ02
    |--Alaus Eschscholtz 1829J02
    |    |--A. infumatus Candeze 1874M86
    |    |--A. lusciosusB14
    |    |--A. obliquus Candeze 1874M86
    |    |--A. oculatusAT01
    |    `--A. zunianus Casey 1893B14
    `--Paracalais Neboiss 1967M85, C96
         |--*P. suboculatus (Candèze 1857) [=Alaus suboculatus; incl. A. variegatus Schwarz 1902]C96
         |--P. albatus (Candèze 1897) [=Alaus albatus]C96
         |--P. darwini (Blackburn 1890) [=Alaus darwini]C96
         |--P. fornicatus Neboiss 1967C96
         |--P. funebris (Candèze 1857) [=Alaus funebris]C96
         |--P. funereus (Candèze 1865) [=Alaus funereus]C96
         |--P. gibboni (Newman 1857) [=Elater gibboni, Alaus gibboni]C96
         |--P. gigas (Candèze 1857) [=Alaus gigas]C96
         |--P. hayekae Neboiss 1967C96
         |--P. lectilis (Candèze 1897) [=Alaus lectilis]C96
         |--P. macleayi (Candèze 1857) [=Alaus macleayi]C96
         |--P. melancholicus (Candèze 1874) [=Alaus melancholicus]C96
         |--P. murinus Neboiss 1967C96
         |--P. nesiotes Neboiss 1967C96
         |--P. prosapius Neboiss 1967C96
         |--P. prosectus (Candèze 1857) [=Alaus prosectus; incl. A. immaculatus Schwarz 1902, A. subsericeus Schwarz 1902]C96
         |--P. pumilis (Candèze 1874) [=Alaus pumilis]C96
         |--P. sericeus (Candèze 1874) [=Alaus sericeus]C96
         |--P. spiciformis Neboiss 1967C96
         |--P. spinicollis (Van Zwaluwenburg 1951) [=Alaus spinicollis]C96
         `--P. victoriae (Schwarz 1902) [=Alaus victoriae]C96

*Type species of generic name indicated


[AT01] Arnett, R. H., Jr & M. C. Thomas (eds) 2001. American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. CRC Press: Boca Raton.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[C96] Calder, A. A. 1996. Click beetles: genera of the Australian Elateridae (Coleoptera). CSIRO Australia: Collingwood.

[FS90] Fleutiaux, E., & A. Sallé. 1890. Liste des coléoptères de la Guadeloupe et descriptions d’espèces nouvelles. Annales de la Société Entomologique de France, 6e série 9: 351–484.

[I92] Imes, R. 1992. The Practical Entomologist. Aurum Press: London.

[JG19] James, D. J., P. T. Green, W. F. Humphreys & J. C. Z. Woinarski. 2019. Endemic species of Christmas Island, Indian Ocean. Records of the Western Australian Museum 34 (2): 55–114.

[J02] Johnson, P. J. 2002. Elateridae Leach 1815. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 160–173. CRC Press.

[KB11] Kundrata, R., & L. Bocak. 2011. The phylogeny and limits of Elateridae (Insecta, Coleoptera): is there a common tendency of click beetles to soft-bodiedness and neoteny? Zoologica Scripta 40: 364–378.

[M86] Macleay, W. 1886. The insects of the Fly River, New Guinea, “Coleoptera”. Proceedings of the Linnean Society of New South Wales, series 2, 1 (1): 136–157.

[M85] Matthews, E. G. 1985. A Guide to the Genera of Beetles of South Australia. Part 4. Polyphaga: Byrrhoidea, Buprestoidea, Dryopoidea, Elateroidea, Cantharoidea, Derodontoidea, and Bostrichoidea. South Australian Museum: Adelaide.

[PS07] Platia, G., & R. Schimmel. 2007. Click beetles of Taiwan collected by the expeditions of the Hungarian Natural History Museum in the years 1995 to 2003 (Coleoptera: Elateridae). Annales Historico-Naturales Musei Nationalis Hungarici 99: 49–91.

Leave a comment

Your email address will not be published. Required fields are marked *