Heterohelix striata, from here.

Belongs within: Globigerinana.

The Heterohelicidae are a group of planktonic Foraminifera known from the Middle Jurassic to the Oligocene (Loeblich & Tappan 1964).

Characters (from Loeblich & Tappan 1964): Early stage trochospiral, planispiral, biserial or triserial, lateral may show serial reductions or proliferations; aperture large, simple and interiomarginal, or terminal in uniserial forms, without internal columellar processes.

|--Gublerina Kikoïne 1948LT64 [GublerininaeM13]
| `--G. ornatissima (Cushman & Church 1929) [=Ventilabrella ornatissima; incl. *G. cuvillieri Kikoïne 1948]LT64
|--Pseudoguembelina Brönnimann & Brown 1953LT64 [PseudogumbelininaeM13]
| `--*P. excolata (Cushman 1926) [=Guembelina excolata]LT64
`--Heterohelicinae [Guembelininae]LT64
| i. s.: Sigalia Reiss 1957LT64
| `--*S. deflaensis (Sigal 1952)LT64, M08b, LT64 (see below for synonymy)
| Racemiguembelina Montanaro Gallitelli 1957LT64
| `--*R. fructicosa (Egger 1899) [=Guembelina fructicosa]LT64
|--Bifarina Parker & Jones 1872C40 [incl. Rectoguembelina Cushman 1932LT64, Tubitextularia Šulc 1929LT64]
| |--*B. saxipara (Ehrenberg 1854) [=Dimorphina saxipara]C40
| |--B. bohemica [=Pseudotextularia bohemica, *Tubitextularia bohemica]LT64
| |--B. fimbriataC40
| |--B. nodosaria White 1929 [incl. *Rectoguembelina cretacea Cushman 1932]LT64
| `--‘Rectoguembelina’ texanaC40
|--Heterohelix Ehrenberg 1841C40, LT61 [=Spiroplecta Ehrenberg 1844LT64; incl. Guembelina Egger 1899 non Kuntze 1895LT64]
| |--*H. americana (Ehrenberg 1844) [=Spiroplecta americana; incl. H. navarroensis]LT64
| |--H. cenomanicaBL79
| |--H. complanataM08b
| |--H. dentata Stenestad 1968FM03
| |--H. globulosa (Ehrenberg 1840) [=Textularia globulosa, *Guembelina globulosa]LT64
| |--H. moremaniI02
| |--H. pulchraM08b
| |--‘Guembelina’ spiniferaC40
| `--H. striataM08b [=Guembelina striataC40]
`--+--Planoglobulina Cushman 1927C40 [incl. Ventilabrella Cushman 1928LT64]
| |--*P. acervulinoides (Egger 1899) [=Guembelina acervulinoides]LT64
| |--‘Ventilabrella’ carseyiC40
| `--*Ventilabrella’ eggeri Cushman 1928C40
`--Pseudotextularia Rzehak 1891C40, LT64 (see below for synonymy)
|--*P. elegans (Rzehak 1891) [=Cuneolina elegans]LT64
|--P. fruticosaC40
|--‘Guembelina’ plummerae Loetterle 1937 [=*Bronnimannella plummerae]LT64
`--P. varians Rzehak 1886C40

Pseudotextularia Rzehak 1891C40, LT64 [incl. Bronnibrownia Montanaro Gallitelli 1955 (n. n.)LT64, Bronnimannella Montanaro Gallitelli 1956LT64]

*Sigalia deflaensis (Sigal 1952)LT64, M08a, LT64 [=Guembelina (Ventilabrella) deflaensisLT64, Gublerina deflaensisLT64]

*Type species of generic name indicated


[BL79] Basov, V. A., B. G. Lopatin, I. S. Gramberg, A. I. Danjushevskaya, V. Ya. Kaban’kov, V. M. Lazurkin & D. K. Patrunov. 1979. Lower Cretaceous lithostratigraphy near Galicia Bank. Initial Reports of the Deep Sea Drilling Project 47: 683–717.

[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3rd ed. Harvard University Press: Cambridge (Massachusetts).

[FM03] Friedrich, O., & K. J. S. Meier. 2003. Stable isotope indication for the cyst formation depth of Campanian/Maastrichtian calcareous dinoflagellates. Micropaleontology 49 (4): 375–380.

[I02] Ibrahim, M. I. A. 2002. Late Albian–Middle Cenomanian palynofacies and palynostratigraphy, Abu Gharadig-5 well, Western Desert, Egypt. Cretaceous Research 23: 775–788.

[LT61] Loeblich, A. R., Jr & H. Tappan. 1961. Suprageneric classification of the Rhizopodea. Journal of Paleontology 35 (2): 245–330.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 2. The Geological Society of America and The University of Kansas Press.

[M08a] McMillan, I. K. 2008a. Aragonitic-walled benthic foraminifera (Epistomina) in the Cretaceous ‘mudbelt’ off southern Africa, and postmortem cross-shelf transport of tests. African Natural History 4: 17–24.

[M08b] McMillan, I. K. 2008b. Reappraisal of foraminiferal assemblages of the Santonian–Campanian Mzamba Formation type section, and their correlation with the stratigraphic succession of the KwaZulu Basin. African Natural History 4: 25–34.

[M13] Mikhalevich, V. I. 2013. New insight into the systematics and evolution of the Foraminifera. Micropaleontology 59 (6): 493–527.

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