Male Heteropachylus inexpectabilis, copyright Pedro H. Martins.

Belongs within: Gonyleptidae.

The Heteropachylinae are a group of gonyleptid harvestmen in which the males have the first two free tergites of the opisthosoma fused to the dorsal scutum (Mendes 2011).

Characters (from Mendes 2011): Scutum length about 5 mm, anterior border of carapace usually with tubercles or granules. Eye mound wider than long, convex, not located directly on anterior margin of carapace, bearing pair of tubercles or granules. Area III with variable armature, mostly with pair of paramedian small tubercles. Males with at least free tergites I and II fused to the dorsal scutum, free tergite II bearing median apophysis (rarely reduced or absent) and armature on corners. Ozopore with V-shaped cut at posterior rim of anterior opening slit. Sternite V double the size of the others in lateral view. Basichelicerite smooth, with ventral granule, bulla well marked; cheliceral hand smooth, with bristles mainly on distal portion. Calcaneus I slightly thicker than astragalus I. Coxae I–II with a pair of dorsal apophyses each, those of coxa II bifurcate. Male coxa IV hypertelic, bearing apical apophysis on the prolateral margin (small spine in females). Male trochanter IV wider than long, with apical curve apophysis on the prolateral margin and acuminate on the retrolateral margin (trochanter narrow in females with apical prolateral spine). Femur IV short, curved in dorsal view, bearing rows of tubercles and apical apophyses in males. Male genitalia with ventral plate subrectangular, slightly narrower at basal portion, distal border concave, without lateral lobes, excavated on dorsolateral surface seeming to be divided into ventral and dorsal layers and ventrally covered with minute setae; blunt small rounded ventral setae and five pairs of dorsal spiniform setae; glans without ventral process, in some cases bearing a dorsal digitiform process. Stylus with subapical small setae.

    |--+--Magnispina Mendes 2011M11
    |  |    `--*M. neptunus Mendes 2011M11
    |  `--Pseudopucrolia Roewer 1912 (see below for synonymy)M11
    |       |--P. incerta (Mello-Leitão 1928) [=Heterogonyleptes incertus, *Melloa incerta, Psudopucrolia (l. c.) incerta]M11
    |       `--+--P. discrepans (Roewer 1943) [=*Tribunosoma discrepans]M11
    |          `--+--P. mutica (Perty 1833)M11, K03 (see below for synonymy)
    |             `--P. rugosa (Roewer 1930) [=*Thaumatoleptes rugosus]M11
    `--Heteropachylus Roewer 1913 [incl. Aesotrinoma Soares 1977, Chavesincola Soares & Soares 1946]M11
         |--H. ramphonotus Mendes 2011M11
         `--+--+--M. crassicalcanei (Soares 1977) [=*Aesotrinoma crassicalcanei, Chavesincola crassicalcanei]M11
            |  `--H. inexpectabilis (Soares & Soares 1946)M11, C-FL13 [=*Chavesincola inexpectabilisK03]
            `--+--H. gracilis Mendes 2011M11
               `--+--*H. spiniger Roewer 1913M11
                  `--H. peracchii Soares & Soares 1974M11, K03

Pseudopucrolia Roewer 1912 [incl. Heterogonyleptes Roewer 1913, Melloa Roewer 1930, Pseudotriaenosoma Mello-Leitão 1927, Thaumatoleptes Roewer 1930, Triaenosoma Roewer 1913, Tribunosoma Roewer 1943]M11

Pseudopucrolia mutica (Perty 1833)M11, K03 [=Eusarcus muticusK03, Eusarchus (l. c.) muticusK03, Gonyleptes muticusK03, *Heterogonyleptes muticusK03; incl. Triaenosoma bahiensis Mello-Leitão 1926K03, *Pseudotriaenosoma pickeli Mello-Leitão 1927K03, *T. singularis Roewer 1913K03, *Pseudopucrolia spinosa Roewer 1912K03]

*Type species of generic name indicated


[C-LF13] Cruz-López, J. A., & O. F. Francke. 2013. On the enigmatic genus Philora: familial assignment and taxonomic revision (Opiliones: Laniatores: Stygnopsidae). Journal of Arachnology 41: 291–305.

[K03] Kury, A. B. 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Ibérica de Aracnología, volumen especial monográfico 1: 1–337.

[M11] Mendes, A. C. 2011. Phylogeny and taxonomic revision of Heteropachylinae (Opiliones: Laniatores: Gonyleptidae). Zoological Journal of the Linnean Society 163: 437–483.

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