Homalenotus quadridentatus, copyright Arp.

Belongs within: Sclerosomatidae.

Homalenotus is a genus of heavily armoured sclerosomatid harvestmen known from central and southern Europe.

Characters (from Martens 1978): Male genital morphology with truncus of penis slender, narrowed continuously from base to glans, the third of which is almost parallel-margined, glans narrow or broad lancet-shaped, sometimes wider than end of truncus, weakly angled dorsad against truncus. Muscles restricted to basal half of truncus. Body of male and female conspicuously dorso-ventrally flattened, anterior margin of cephalothorax drawn out medially into 2 superimposed, horizontal processes, the upper strikingly large and only this one visible dorsally. Metapeltidium without or with two median humps, opisthosomal areas I–IV with one pair of median humps, laterally often with another pair. Area V extended in 4 horizontally protruding processes. Chelicerae almost completely concealed dorsally. Supracheliceral lamellae weakly developed, only visible ventrally after removal of the chelicerae. Tarsal setation of pedipalp strong, comb-like. Sternites I-VI each with indistinctly differentiated lateral sclerite plate in region of pleura. Corona analis present.

<==Homalenotus Koch 1839G59 (see below for synonymy)
    |--*H. quadridentatus (Cuvier 1795)C92 (see below for synonymy)
    |--H. armatus (Roewer 1915)G59 [=*Parasclerosoma armatumC92]
    |--H. buchneri Schenkel 1936G59 (see below for synonymy)
    |--H. coriaceus (Simon 1879)G59 (see below for synonymy)
    |--H. graecus Roewer 1957G59
    |--H. laranderas Grasshoff 1959G59
    |--‘Phalangium’ mamillatum Gervais 1844 (n. d.) [=Sclerosoma mamillatum]R23
    |--H. monoceros Koch 1839G59 [=Sclerosoma monocerosD04, H. monocerus (l. c.)D04]
    |--‘Phalangium’ oraniense Lucas 1847 [=Sclerosoma oraniense]R23
    `--H. remyi (Roewer 1957) [=Parasclerosoma remyi]G59

Homalenotus Koch 1839G59 [=Homalonotus Agassiz 1846 nec Koenig 1825 nec Agassiz 1846C92, Sclerosoma Lucas 1858C92; incl. Parasclerosoma Roewer 1915G59]

Homalenotus buchneri Schenkel 1936G59 [=Parasclerosoma buchneriR57; incl. P. lusitanicum Roewer 1923 non Sclerosoma lusitanicum Kulczynski 1909G59, Homalenotus maroccanus Roewer 1957PWF22, H. roeweri Kraus 1959G59]

Homalenotus coriaceus (Simon 1879)G59 [=Sclerosoma coriaceumG59; incl. H. hispanum Koch 1869 (n. n.?)R23, S. hispanumR23, S. lusitanicum Kulczynski 1909G59, Homalenotus lusitanicusG59]

*Homalenotus quadridentatus (Cuvier 1795)C92 [=Phalangium quadridentatumG59, *Sclerosoma quadridentatumC92; incl. Homalenotus bispinosus Lucas 1856D04, S. bispinosumD04, Parasclerosoma perssoni brunnea Schenkel 1938D04, Pa. perssoni Schenkel 1938D04, Homalenotus romanus Koch 1869D04, Sclerosoma romanumD04, Phalangium spinosum Latreille 1798 non Bosc 1792D04]

*Type species of generic name indicated


[C92] Crawford, R. L. 1992. Catalogue of the genera and type species of the harvestman superfamily Phalangioidea (Arachnida). Burke Museum Contributions in Anthropology and Natural History 8: 1–60.

[D04] Delfosse, E. 2004. Catalogue préliminaire des Opilions de France métropolitaine (Arachnida Opiliones). Bulletin de Phyllie 20: 34–58.

[G59] Grasshoff, M. 1959. Über Homalenotus und Parasclerosoma. (Arach., Opiliones-Palpatores). Senckenbergiana Biologica 40 (5–6): 283–288.

[PWF22] Prieto, C. E., H. Wijnhoven & J. B. Febrer. 2022. New harvestmen (Arachnida: Opiliones) from Menorca Island. Revista Ibérica de Aracnología 40: 145–157.

[R23] Roewer, C.-F. 1923. Die Weberknechte der Erde: Systematische Bearbeitung der bisher bekannnten Opiliones. Gustav Fischer: Jena.

[R57] Roewer, C. F. 1957. Über Oligolophinae, Caddoinae, Sclerosomatinae, Leiobuninae, Neopilioninae und Leptobuninae (Phalangiidae, Opiliones Palpatores). Senckenbergiana Biologica 38 (5–6): 323–358.

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