Aggregation of Humerobates rostrolamellatus, copyright Theodoor Heijerman.

Belongs within: Ceratozetoidea.

Humerobates is a cosmopolitan genus of oribatid mites in which the pteromorphs may be movable or immovable, and the notogastral setae are represented by alveoli only (Balogh & Balogh 1992). The subgenus Cordylobates is represented by a single species H. (C.) fragilis from Ascension Island in the South Atlantic.

Characters (from Balogh & Balogh 1992): Prodorsum with lamellae present, translamella absent. Notogaster with ten pairs of setal alveoli only; four pairs of areae porosae; pteromorphs well-developed, movable or immovable. Six pairs of genital setae; one pair of aggenital setae; two pairs of anal setae; three pairs of adanal setae. Legs tridactyle.

<==Humerobates Sellnick 1928S04
    |--H. (Humerobates) (see below for synonymy)S04
    |    |--H. (H.) rostrolamellatus Grandjean 1936S04 (see below for synonymy)
    |    |    |--H. r. rostrolamellatusS04
    |    |    |--H. r. giganteus Willmann 1939 [incl. H. fungorum giganteus]S04
    |    |    |--H. r. guadarramicus Pérez-Íñigo 1972 [incl. Chamobates magnus Mihelčič 1957 (n. d.)]S04
    |    |    `--H. r. mauritius Jacot 1936 [=H. humeralis mauritius]S04
    |    |--H. (H.) africanus (Mahunka 1984) [=Baloghobates africanus]S04
    |    |--H. (H.) cornutus (Sellnick 1959)S04 [=*Trihumerozetes cornutusBB92]
    |    |--H. (H.) crozetensis (Richters 1908)S04 (see below for synonymy)
    |    |--H. (H.) flechtmanni Pérez-Íñigo & Pérez-Íñigo 1993S04
    |    |--H. (H.) fungorum (Linnaeus 1758) (n. d.) [=Acarus fungorum]S04
    |    |--‘Antarctozetes’ intermedius (Hammer 1967)L85
    |    |--‘Antarctozetes’ longicaulis (Hammer 1967)L85
    |    |--‘Antarctozetes’ luteus (Hammer 1967)L85
    |    |--H. (H.) nudus (Hammer 1967)S04 [=*Baloghobates nudusCH98]
    |    |--H. (H.) papuanus Balogh 1970S04
    |    |--H. (H.) parvoglobosus (Hammer 1967) [=Baloghobates parvoglobosus]S04
    |    |--H. (H.) perkinsi Jacot 1934S04 [=H. humeralis perkinsiS04, H. rostrolamellatus perkinsiSN98]
    |    |--H. (H.) pomboi Pérez-Íñigo 1992S04
    |    |--H. (H.) setosus Behan-Pelletier & Mahunka 1993S04
    |    |--H. (H.) taiwanensis Tseng 1984S04
    |    |--H. (H.) varius Ohkubo 1982S04
    |    `--H. (H.) zumpti Sellnick 1957S04
    `--H. (Cordylobates Luxton 1995)S04
         `--H. (*C.) fragilis (Luxton 1995) [=*Cordylobates fragilis]S04

Humerobates Sellnick 1928 (Humerobates) [incl. Antarctozetes Balogh 1961, Baloghobates Hammer 1967, Banksinus Jacot 1938, Jeannelia Dalenius 1958 non Raffray 1913, Trihumerozetes Sellnick 1959]S04

Humerobates (Humerobates) crozetensis (Richters 1908)S04 [=Oribata crozetensisS04, *Antarctozetes crozetensisCH98, *Jeannelia crozetensisCH98]

Humerobates (Humerobates) rostrolamellatus Grandjean 1936S04 [=Banksinus arboreus rostrolamellatusJ38; incl. Oribata arborea Banks 1895 (n. d.)S04, *Banksinus arboreusJ38, Humerobates humeralis arboreaS59, Sphaerobates dentatus Mihelčič 1957S04, Notaspis humeralis Sellnick 1928 non Hermann 1804S04, *Humerobates humeralisS04, Murcia rubra Koch 1841 (n. d.)S04, Oribata rubraM98]

*Type species of generic name indicated


[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[CH98] Colloff, M. J., & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[J38] Jacot, A. P. 1938. Some new western North Carolina moss-mites. Proceedings of the Entomological Society of Washington 40 (1): 10–15.

[L85] Luxton, M. 1985. Cryptostigmata (Arachnida: Acari)—a concise review. Fauna of New Zealand 7: 1–106.

[M98] Michael, A. D. 1898. Oribatidae. In: H. Lohmann (ed.) Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen vol. 3. Acarina pp. 1–93. R. Friedländer und Sohn: Berlin.

[S59] Sellnick, M. 1959. Acarina from southeastern Polynesia—II (Oribatidae). Occasional Papers of the Bernice P. Bishop Museum 22 (9): 109–152.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

[SN98] Swift, S. F., & R. A. Norton. 1998. Preliminary report on oribatid mite (Acari: Oribatida) diversity in the Hawaiian Islands. Bishop Museum Occasional Papers 57: 1–44.

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