Hyles

Bedstraw hawkmoth Hyles gallii, copyright Tony Morris.

Belongs within: Macroglossini.

Hyles is a cosmopolitan genus of hawkmoths with fore wings typically marked by a pale median stripe (Pittaway 1993).

Characters (from Pittaway 1993): Imago with fore wing entire with apex pointed. Fore wing pattern usually consisting of a pale cream or white, oblique median stripe. Antenna distally incrassate in both sexes, strongly clavate in female. Pilifer with long setae medially, short setae laterally. Eye lashed. Labial palpus smoothly scaled with no erect hairs, concealing base of proboscis; scaling at apex of first segment not arranged in a regular border on inner surface; second segment without apical tuft on inner side. Abdomen conically pointed, with strong dorsal spines usually arranged in three rows. External spines of fore tarsus more or less prolonged, always longer than respective spines on inner side of tarsus; comb of mid and hind tarsus vestigial, the spines not much prolonged. First segment of hind tarsus shorter than tibia. Inner distal spur of hind tibia more than twice as long as outer and about half tibial length. Pulvillus absent or vestigial. Genitalia in male with valva broadly sole-shaped; friction scales usually numerous and small. Sacculus ending in a thin, more or less curved, simple, tapering process. Aedeagus with dorsal apical edge incrassate, dentate, produced at left side into short process. Ovum small, almost spherical, pale green or blue-green. Larva typically sphingiform; thorax slightly tapering anteriorly. Usually with conspicuous dorso-lateral line of eye-spots. Pupa with proboscis fused with body, not projecting anteriorly, not obviously carinate. Light tan, smooth, covered with short, fine dark lines. Cremaster large, sometimes bent ventrally, with apical pair of spines. In a fairly substantial cocoon on ground.

<==Hyles Hübner 1819 [incl. Celerio Agassiz 1846, Hawaiina Tutt 1903, Turneria Tutt 1903 non Forel 1895]P93
    |--*H. euphorbiae (Linnaeus 1758) [=Sphinx euphorbiae]P93
    |    |--H. e. euphorbiae (see below for synonymy)P93
    |    |--H. e. conspicua (Rothschild & Jordan 1903) [=Celerio euphorbiae conspicua]P93
    |    `--H. e. robertsi (Butler 1880) (see below for synonymy)P93
    |--H. celerioDS73
    |--H. centralasiae (Staudinger 1887) [=Deilephila euphorbiae var. centralasiae]P93
    |    |--H. c. centralasiae [incl. Celerio centralasiae transcaspica Bang-Haas 1936]P93
    |    `--H. c. siehei (Püngeler 1903) [=Deilephila siehei]P93
    |--H. chamyla (Denso 1913) [=Celerio hippophaes chamyla; incl. C. chamyla apocyni Shchetkin 1956]P93
    |--H. costata (Nordmann 1841)P93
    |    |--H. c. costataP93
    |    `--H. c. sinensis (Closs 1917) [=Celerio euphorbiae sinensis]P93
    |--H. dahlii (Geyer in Hübner 1827) [=Sphinx dahlii; incl. Celerio euphorbiae balearica Rebel 1926]P93
    |--H. gallii (Rottemburg 1775) [=Sphinx gallii, S. galii Denis & Schiffermüller 1775]P93
    |    |--H. g. gallii (see below for synonymy)P93
    |    |--H. g. intermedia (Kirby 1837)P93
    |    |--H. g. nepalensis (Daniel 1960)P93
    |    `--H. g. tibetanica (Eichler 1971)P93
    |--H. hippophaes (Esper 1793) [=Sphinx hippophaes]P93
    |    |--H. h. hippophaes [incl. Celerio hippophaes kiortsii Koutsaftikis 1974]P93
    |    |--H. h. bienerti (Staudinger 1874) (see below for synonymy)P93
    |    `--H. h. caucasica (Denso 1913) [=Celerio hippophaes caucasica]P93
    |--H. lineata (Fabricius 1775)P93 [=Celerio lineataC70]
    |    |--H. l. lineataPP72
    |    `--‘Celerio’ l. florilegaPP72
    |--H. livornica (Esper 1779) [=Sphinx livornica]P93
    |    |--H. l. livornica [incl. Phinx koechlini Fuessly 1781, Celerio lineata saharae Stauder 1921]P93
    |    |--H. l. livornicoides (Lucas 1891)P93 [=Celerio lineata livornicoidesC70, H. lineata livornicoidesMC13]
    |    |--H. l. malgassica (Denso 1944)P93
    |    `--H. l. tatsienluica Oberthür 1916P93
    |--H. nervosa (Rothschild & Jordan 1903) [=Celerio euphorbiae nervosa]P93
    |--H. nicaea (de Prunner 1798) [=Sphinx nicaea]P93
    |    |--H. n. nicaea [incl. H. nicaea nicaea f. albina, Sphinx cyparissiae Hübner 1805, H. nicaea nicaea f. rubida]P93
    |    |--H. n. castissima (Austaut 1883) [=Deilephila nicaea var. castissima]P93
    |    |--H. n. crimaea (Bang-Haas 1906) [=Deilephila nicaea crimaea; incl. Celerio nicaea var. orientalis Austaut 1905]P93
    |    `--H. n. libanotica (Gehlen in Seitz 1932) (see below for synonymy)P93
    |--H. salangensis (Ebert 1969) [=Celerio salangensis]P93
    |--H. tithymali (Boisduval 1832) [=Deilephila tithymali]P93
    |    |--H. t. tithymali [incl. D. calverleyi Grote & Robinson 1865]P93
    |    |--H. t. deserticola (Staudinger in Staudinger & Rebel 1901) (see below for synonymy)P93
    |    |--H. t. gecki de Freina 1991 [=H. euphorbiae gecki]P93
    |    |--H. t. himyarensis Meerman 1988P93
    |    `--H. t. mauretanica (Staudinger in Staudinger & Wocke 1871) [=Deilephila mauretanica, H. euphorbiae mauretanica]P93
    |--H. vespertilio (Esper 1779) [=Sphinx vespertilio; incl. H. vespertilio f. flava, H. vespertilio f. salmonea]P93
    `--H. zygophylli (Ochsenheimer 1808) [=Sphinx zygophylli]P93
         |--H. z. zygophylli [incl. Celerio zygophylli jaxartis Froreich 1938]P93
         `--H. z. xanthoxyli (Derzhavets 1977)P93

Hyles euphorbiae euphorbiae (Linnaeus 1758) [incl. H. euphorbiae euphorbiae f. brunnescens, Celerio euphorbiae dolomiticola Stauder 1930, Sphinx esulae Hufnagel 1766, Deilephila esulae Boisduval 1834 non S. esulae Hufnagel 1766, Celerio euphorbiae etrusca Verity 1911, C. euphorbiae filapjewi Bang-Haas 1936, Deilephila euphorbiae grentzenbergi Staudinger 1885, Hyles euphorbiae euphorbiae f. lafitolei, H. euphorbiae lucida Derzhavets 1980, Deilephila paralias Nickerl 1837, H. euphorbiae euphorbiae f. restricta, Celerio euphorbiae rothschildi Stauder 1928, C. euphorbiae strasillai Stauder 1921, C. euphorbiae subiacensis Dannehl 1929, C. euphorbiae vandalusica Rebel 1910]P93

Hyles euphorbiae robertsi (Butler 1880) [=Deilephila robertsi; incl. H. robertsi orientalis Ebert 1969 non Celerio nicaea orientalis Austraut 1905, D. peplidis Christoph 1894]P93

Hyles gallii gallii (Rottemburg 1775) [incl. Celerio gallii chishimana Matsumura 1929, Hyles gallii gallii f. grisea, Hyles gallii gallii f. pallida, C. gallii sachaliensis Matsumura 1929]P93

Hyles hippophes bienerti (Staudinger 1874) [=Deilephila bienerti; incl. Celerio hippophaes anatolica Rebel 1933, C. hippophaes bucharana Sheljuzhko 1933, Deilephila insidiosa Erschoff in Fedtshenko 1874, C. hyppophaes malatiatus Gehlen 1934, C. hippophaes ornatus Gehlen 1930, C. hippophaes shugnana Sheljuzhko 1933, C. hippophaes transcaucasica Gehlen in Seitz 1932]P93

Hyles nicaea libanotica (Gehlen in Seitz 1932) [=Celerio nicaea libanotica; incl. C. nicaea var. sheljuzkoi Dublitzky 1928, Hyles nicaea sheljuzkoi]P93

Hyles tithymali deserticola (Staudinger in Staudinger & Rebel 1901) [=Deilephila mauretanica deserticola, H. euphorbiae deserticola, C. deserticola saharae Günther 1939 non C. lineata saharae Stauder 1921]P93

*Type species of generic name indicated

References

[C70] Common, I. F. B. 1970. Lepidoptera (moths and butterflies). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 765–866. Melbourne University Press.

[DS73] Dickens, M., & E. Storey. 1973. The World of Moths. Macmillan Publishing Co., Inc.: New York.

[MC13] Majer, J. D., S. K. Callan, K. Edwards, N. R. Gunawardene & C. K. Taylor. 2013. Baseline survey of the terrestrial invertebrate fauna of Barrow Island. Records of the Western Australian Museum Supplement 83: 13–112.

[PP72] Parkin, P., D. T. Parkin, A. W. Ewing & H. A. Ford. 1972. A report on the arthropods collected by the Edinburgh University Galapagos Islands expedition, 1968. Pan-Pacific Entomologist 48 (2): 100–107.

[P93] Pittaway, A. R. 1993. The Hawkmoths of the Western Palaearctic. Harley Books: Colchester.

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