Stephen’s rocket frog Anomaloglossus stepheni, copyright Diogo Luiz.

Belongs within: Neobatrachia.
Contains: Alsodidae, Batrachylidae, Hylodidae, Telmatobius, Ceratophryidae, Hemiphractidae, Hylidae, Phyllomedusidae, Litoria, Pelodryadinae, Dendrobatidae, Aromobatidae, Brachycephalidae, Craugastoridae, Strabomantidae, Eleutherodactylus, Bufonidae, Leptodactylidae, Centrolenidae, Odontophrynidae.

The Hyloidea are a diverse clade of frogs united by molecular data.

<==Hyloidea [Agastorophrynia, Dendrobatoidea, Hesticobatrachia]PW11
|--Baurubatrachus pricei Báez & Perí 1987DMH16, R00
`--+--Rhinodermatidae [Rhinodermina]FB17
| |--Insuetophrynus Barrio 1970PW11, FG06
| | `--I. acarpicusPW11
| `--Rhinoderma Duméril & Bibron 1841FG06
| |--R. darwiniiPW11
| `--R. rufumFG06
| `--+--BatrachylidaeFB17
| `--HylodidaeFB17
`--+--Telmatobiidae [Batrachophrynidae]FB17
| |--TelmatobiusPW11
| |--Neoprocoela Schaeffer 1949B00
| | `--*N. edentatus Schaeffer 1949B00
| |--Batrachophrynus Peters 1873PW11, FG06
| | |--B. brachydactylusFG06
| | `--B. macrostomusFB17
| `--Caudiverbera Laurenti 1768FG06 (see below for synonymy)
| |--C. caudiverbera (see below for synonymy)B00
| `--*Eophractus’ casamayorensis Schaeffer 1949B00
| `--+--HemiphractidaeFB17
| `--ArboranaeDMH16
| |--HylidaeFB17
| `--+--PhyllomedusidaeDMH16
| `--PelodryadidaeDMH16
| | i. s.: Australobatrachus iliusDMH16
| |--LitoriaFG06
| `--PelodryadinaeFB17
| `--+--Rheobates palmatusPW11
| `--+--AromobatidaeGM19
| `--AnomaloglossusPW11
| |--+--A. baeobatrachusPW11
| | `--A. stepheniPW11
| `--+--A. tepuyensisPW11
| `--+--A. degranvilleiPW11
| `--+--A. praderioiPW11
| `--+--A. beebeiPW11
| `--A. roraimaPW11
`--+--+--Ceuthomantis [Ceuthomantidae]PW11
| | `--C. smaragdinusPW11
| `--+--BrachycephalidaePW11
| `--+--+--CraugastoridaePW11
| | `--StrabomantidaeFB17
| `--EleutherodactylidaeFB17
| |--EleutherodactylusFG06
| `--PhyzelaphryninaePW11
| |--Phyzelaphryne Heyer 1977FG06
| | `--P. miriamaePW11
| `--Adelophryne Hoogmoed & Lescure 1984PW11, FG06
| `--A. gutturosaPW11
| `--+--CentrolenidaePW11
| `--Allophryne Gaige 1926FG06 [AllophrynidaePW11, Allophryninae]
| `--A. ruthveniPW11
`--Cycloramphidae [Cyclorhamphina, Cycloramphinae, Grypiscina, Grypiscini]FB17
|--Rupirana Heyer 1999FG06
|--Crossodactylodes Cochran 1938FG06
|--Zachaenus Cope 1866FG06
|--Grypiscus Cope 1867FG06
|--Thoropa Cope 1865PW11, FG06 [Thoropidae]
| |--T. miliarisPW11
| `--T. taophoraPW11
`--Cycloramphus Tschudi 1838FG06
|--C. acangatanPW11
|--C. boraceiensisPW11
`--C. stejnegeriFG06

Caudiverbera Laurenti 1768FG06 [incl. Eophractus Schaeffer 1949B00, Gigantobatrachus Casamiquela 1958B00, Teracophrys Ameghino 1901B00]

Caudiverbera caudiverbera [incl. Calyptocephalella canqueli Schaeffer 1949, *Gigantobatrachus parodii Casamiquela 1958]B00

*Type species of generic name indicated


[B00] Báez, A. M. 2000. Tertiary anurans from South America. In: Heatwole, H., & R. L. Carroll (eds) Amphibian Biology vol. 4. Palaeontology. The evolutionary history of amphibians pp. 1388–1401. Surrey Beatty & Sons.

[DMH16] Duellman, W. E., A. B. Marion & S. B. Hedges. 2016. Phylogenetics, classification, and biogeography of the treefrogs (Amphibia: Anura: Arboranae). Zootaxa 4104 (1): 1–109.

[FB17] Feng, Y.-J., D. C. Blackburn, D. Liang, D. M. Hillis, D. B. Wake, D. C. Cannatella & P. Zhang. 2017. Phylogenomics reveals rapid, simultaneous diversification of three major clades of Gondwanan frogs at the Cretaceous–Paleogene boundary. Proceedings of the National Academy of Sciences of the USA 114 (29): E5864–E5870.

[FG06] Frost, D. R., T. Grant, J. Faivovich, R. H. Bain, A. Haas, C. F. B. Haddad, R. O. de Sá, A. Channing, M. Wilkinson, S. C. Donnellan, C. J. Raxworthy, J. A. Campbell, B. L. Blotto, P. Moler, R. C. Drewes, R. A. Nussbaum, J. D. Lynch, D. M. Green & W. C. Wheeler. 2006. The amphibian tree of life. Bulletin of the American Museum of Natural History 297: 1–370.

[GM19] Guillory, W. X., M. R. Muell, K. Summers & J. L. Brown. 2019. Phylogenomic reconstruction of the Neotropical poison frogs (Dendrobatidae) and their conservation. Diversity 11: 126.

[PW11] Pyron, R. A., & J. J. Wiens. 2011. A large-scale phylogeny of Amphibia including over 2800 species, and a revised classification of extant frogs, salamanders, and caecilians. Molecular Phylogenetics and Evolution 61: 543–583.

[R00] Roček, Z. 2000. Mesozoic anurans. In: Heatwole, H., & R. L. Carroll (eds) Amphibian Biology vol. 4. Palaeontology. The evolutionary history of amphibians pp. 1295–1331. Surrey Beatty & Sons.

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