Iguanodontia

Reconstruction of Iguanodon atherfieldensis, copyright Steveoc 86.

Belongs within: Ornithopoda.
Contains: Hadrosauroidea.

The Iguanodontia are a diverse group of bipedal and facultatively quadrupedal herbivorous dinosaurs known from the Late Jurassic and Cretaceous, recognised by Norman (2004) as the clade of all taxa closer to Edmontosaurus than Thescelosaurus.

Characters (from Norman 2004): Premaxilla with transversely expanded and edentulous margin; predentary with smoothly convex rostral margin in dorsal aspect; dentary ramus deep, with parallel dorsal and ventral borders; sternal rib ossification absent; one phalanx lost on manus digit III; prepubic process compressed, blade-shaped.

<==Iguanodontia [Iguanodontidae]
    |--Tenontosaurus Ostrom 1970N04
    |    |--T. dossi Winkler, Murry & Jacobs 1997N04
    |    `--T. tilletti Ostrom 1970N04
    `--+--Muttaburrasaurus Bartholomai & Molnar 1981N04
       |    `--M. langdoni Bartholomai & Molnar 1981N04
       `--EuiguanodontiaCPN07
            |--Gasparinisaura Coria & Salgado 1996CPN07, NS04
            |    `--G. cincosaltensis Coria & Salgado 1996NS04
            `--+--ElasmariaCPN07
               |    |--Talenkauen santacrucensisCPN07
               |    `--Macrogryphosaurus Calvo, Porfiri & Novas 2007CPN07
               |         `--*M. gondwanicus Calvo, Porfiri & Novas 2007CPN07
               `--+--Anabisetia Coria & Calvo 1998CPN07, NS04
                  |    `--A. saldiviai Coria & Calvo 2002NS04
                  `--DryomorphaN04
                       |  i. s.: Callovosaurus Galton 1980 (n. d.)N85, N04
                       |           `--*C. leedsi (Lydekker 1889) [=Camptosaurus leedsi]N04
                       |         Kangnasaurus coetzeei Haughton 1915 (n. d.)N85, N04
                       |--DryosauridaeN04
                       |    |--Planicoxa DiCroce & Carpenter 2001N04
                       |    |    `--P. venenica DiCroce & Carpenter 2001N04
                       |    |--Dryosaurus Marsh 1894 [incl. Dysalotosaurus Virchow 1919]N04
                       |    |    |--D. altus (Marsh 1878) [=Laosaurus altus]N04
                       |    |    `--D. lettowvorbecki (Virchow 1919) [=Dysalotosaurus lettowvorbecki]N04
                       |    `--Valdosaurus Galton 1977N04
                       |         |--V. canaliculatus (Galton 1975) (see below for synonymy)N04
                       |         `--V. nigeriensis Galton & Taquet 1982N04
                       `--AnkylopollexiaN04
                            |  i. s.: Draconyx Mateus & Antunes 2001N04
                            |           `--D. loureiroi Mateus & Antunes 2001N04
                            |         Theiophytalia kerriP10
                            |--Camptosaurus Marsh 1885N04 (see below for synonymy)
                            |    |--C. amplus (Marsh 1879) [=Camptonotus amplus]N04
                            |    |--C. aphanoecetesP10
                            |    |--C. depressus Gilmore 1909N04
                            |    |--C. dispar (Marsh 1879) (see below for synonymy)N04
                            |    |--C. hoggi (Owen 1874) [=Iguanodon hoggi]N04
                            |    `--C. prestwichii (Hulke 1880) [=Iguanodon prestwichii]N04
                            `--StyracosternaN04
                                 |--Lurdusaurus Taquet & Russell 1999 [incl. Gravisaurus Chabli 1988]N04
                                 |    `--L. arenatus Taquet & Russell 1999 [incl. Gravisaurus tenerensis Chabli 1988]N04
                                 |--Equijubus You, Luo et al. 2003N04
                                 |    `--E. normani You, Luo et al. 2003N04
                                 `--Iguanodontoidea [Hadrosauriformes]N04
                                      |  i. s.: Fukuisaurus Kobayashi & Azuma 2003N04
                                      |           `--F. tetoriensius Kobayashi & Azuma 2003N04
                                      |         Lanzhousaurus magnidensP10
                                      |         Dollodon bambingiP10
                                      |         Cedrorestes crichtoniP10
                                      |--HadrosauroideaN04
                                      `--Iguanodon Mantell 1825 (see below for synonymy)N04
                                           |--*I. anglicus Holl 1829WU03, N04 (see below for synonymy)
                                           |--I. atherfieldensis Hooley 1924N04 (see below for synonymy)
                                           |--I. bernissartensis Boulenger 1881 [incl. I. seelyi Hulke 1882]N04
                                           |--I. dawsoni Lydekker 1888N04
                                           |--I. exogirarus Fritsch 1878 (n. d.)N04
                                           |--I. fittoni Lydekker 1889 [incl. I. hollingtoniensis Lydekker 1889]N04
                                           |--I. lakotaensis Weishampel & Bjork 1989N04 [=Dakotadon lakotaensisP10]
                                           |--I. orientalis Rozhdestvensky 1952B93
                                           `--I. ottingeri Galton & Jensen 1979 (n. d.)N04
Iguanodontia incertae sedis:
  Albisaurus scutifer Fritsch 1905 (n. d.)N04
  Loncosaurus argentinus Ameghino 1898 (n. d.)N04
  Tichosteus (n. d.)N04
    |--T. aequifacies Cope 1877 (n. d.)N04
    `--T. lucasanus Cope 1877 (n. d.)N04
  RhabdodontidaeWJ03
    |--Rhabdodon Matheron 1869 [incl. Mochlodon Seeley 1881]N04
    |    |--R. priscus Matheron 1869 (see below for synonymy)N04
    |    `--R. septimanicus Buffetaut & le Loeuff 1991N04
    `--Zalmoxes Weishampel, Jianu et al. 2003N04
         |--*Z. robustus (Nopcsa 1902)WJ03 (see below for synonymy)
         `--Z. shqiperorum Weishampel, Jianu et al. 2003N04

Camptosaurus Marsh 1885N04 [incl. Brachyrophus Cope 1878N04, Camptonotus Marsh 1879M96, Cumnoria Seeley 1999N04, Symphyrophus Cope 1878N04; Camptosauridae]

Camptosaurus dispar (Marsh 1879) [=Camptonotus dispar; incl. Brachyrophus altarkansanus Cope 1878, Camptosaurus browni Gilmore 1909, Camptosaurus medius Marsh 1894, Symphyrophus musculosus Cope 1878, Camptosaurus nanus Marsh 1894]N04

Iguanodon Mantell 1825 [incl. Heterosaurus Cornuel 1850, Iguanosaurus Ritgen 1828]N04

*Iguanodon anglicus Holl 1829WU03, N04 [incl. Cetiosaurus brevis Owen 1842N04, Streptospondylus major Owen 1842N04]

Iguanodon atherfieldensis Hooley 1924N04 [=Mantellisaurus atherfieldensisP10; incl. Sphenospondylus gracilis Lydekker 1888N04, I. mantelli Mayer 1832N04, Heterosaurus neocomiensis Cornuel 1850N04, Vectisaurus valdensis Hulke 1879N04]

Rhabdodon priscus Matheron 1869 [incl. Oligomerus adelus Seeley 1881, Ornithomerus gracilis Seeley 1881, Iguanodon suessi Bunzel 1871, Mochlodon suessi]N04

Valdosaurus canaliculatus (Galton 1975) [=Dryosaurus canaliculatus; incl. Camptosaurus valdensis Lydekker 1889]N04

*Zalmoxes robustus (Nopcsa 1902)WJ03 [=Mochlodon robustusN04, Mochlodon robustumWJ03, Rhabdodon robustusWJ03; incl. Onychosaurus hungaricus Nopcsa 1902N04, Camptosaurus inkeyi Nopcsa 1900N04]

*Type species of generic name indicated

References

[B93] Benton, M. J. 1993. Reptilia. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 681–715. Chapman & Hall: London.

[CPN07] Calvo, J. O., J. D. Porfiri & F. E. Novas. 2007. Discovery of a new ornithopod dinosaur from the Portezuelo Formation (Upper Cretaceous), Neuquén, Patagonia, Argentina. Arquivos do Museu Nacional, Rio de Janeiro 65 (4): 471–483.

[M96] Marsh, O. C. 1896. The Dinosaurs of North America: Washington.

[N85] Norman, D. 1985. The Illustrated Encyclopedia of Dinosaurs. Salamander Books: London.

[N04] Norman, D. B. 2004. Basal Iguanodontia. In: Weishampel, D. B., P. Dodson & H. Osmólska (eds) The Dinosauria 2nd ed. pp. 413–437. University of California Press: Berkeley.

[NS04] Norman, D. B., H.-D. Sues, L. M. Witmer & R. A. Coria. 2004. Basal Ornithopoda. In: Weishampel, D. B., P. Dodson & H. Osmólska (eds) The Dinosauria 2nd ed. pp. 393–412. University of California Press: Berkeley.

[P10] Paul, G. S. 2010. Dinosaurs: A Field Guide. A & C Black.

[WJ03] Weishampel, D. B., C.-M. Jianu, Z. Csiki & D. B. Norman. 2003. Osteology and phylogeny of Zalmoxes (n. g.), an unusual euornithopod dinosaur from the latest Cretaceous of Romania. Journal of Systematic Palaeontology 1: 65–123.

[WU03] Wilson, J. A., & P. Upchurch. 2003. A revision of Titanosaurus Lydekker (Dinosauria—Sauropoda), the first dinosaur genus with a ‘Gondwanan’ distribution. Journal of Systematic Palaeontology 1 (3): 125–160.

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