
Belongs within: Percomorpharia.
Contains: Choerodon, Cheilinus, Xanothon, Lepidaplois, Julidinae, Labrus, Scarus.
The Labridae, wrasses, are colourful warm-water marine fishes characterised by sharp, heavy teeth at the front of the mouth in conjunction with crushing pharyngeal jaws (Bond 1996).
<==Labridae [Callyodontidae, Coridae, Scaridae] |--+--HarpinaeR13 | | |--Lachnolaimus maximusND13 | | |--Harpe rufaR13, F15 | | |--PimelometoponR13 | | |--Semicossyphus pulcherR13, CH97 | | `--TrochocopusR13 | `--+--Haletta semifasciataND13 | `--+--Clepticus [Clepticinae]ND13 | | `--C. parraeND13 | `--BodianusND13 | |--B. anthioides (Bennett 1831) [=Crenilabrus anthioides]M58 | |--B. axillaris (Bennett 1832)MM09 | |--B. bilunulatus (Lacépède 1802) [=Labrus bilunulatus, Cossyphus bilunulatus]M58 | |--B. diana (Lacépède 1802) [=Labrus diana]M58 | |--B. mesothorax (Schneider 1801) [=Labrus mesothorax]M58 | |--B. oxycephalus (Bleeker 1862) [=Cossyphus oxycephalus]M58 | |--B. perditioG75 | `--B. rufusND13 `--+--JulidinaeND13 `--+--LabrinaeR13 | |--LabrusL58 | |--Ctenolabrus rupestrisR13, B79 | |--Tautogolabrus adspersus (Walbaum 1792)ND13, M02 | |--Tautoga ornitis (Linnaeus 1758)ND13, M02 | |--AcantholabrusR13 | |--CentrolabrusR13 | `--CrenilabrusR13 | |--C. coeruleusD56 | |--C. griseusD56 | |--C. melopsB79 | `--C. pavoD56 `--+--+--Epibulus Cuvier 1814ND13, S57 [Epibulinae] | | |--E. brevisND13 | | `--E. insidiator (Pallas 1770)S57 [=Sparus insidiatorM58; incl. E. striatus Day 1876S57] | `--OxycheilinusND13 | |--O. celebicus (Bleeker 1853)MM09 | |--O. diagrammus (Lacépède 1801)MM09 | `--O. unifasciatus (Streets 1877)MM09 `--+--CalotomusND13 | |--C. carolinusND13 | |--C. genistriatusJR10 | |--C. moluccensisJR10 | `--C. vaigiensisJR10 `--+--CryptotomusND13 | |--C. beryllinusF16 | |--C. roseus [incl. C. crassiceps Bean 1906]F15 | |--C. spinidens (Quoy & Gaimard 1824) (see below for synonymy)M58 | `--C. ustus [=Callyodon ustus]F16 `--+--Cetoscarus Smith 1956ND13, S59 | |--*C. pulchellus (Rüppell 1835)S59 (see below for synonymy) | `--C. bicolor (Rüppell 1828)S59 (see below for synonymy) `--+--ScarusND13 `--Chlorurus Swainson 1839ND13, S59 [incl. Pseudoscarus Bleeker 1861S59] |--*C. gibbus (Ruppell 1828) [=Scarus gibbus]S59 |--‘Pseudoscarus’ guacamaiaF15 |--C. microrhinos (Bleeker 1854)S59 (see below for synonymy) |--‘Pseudoscarus’ rostratus Gunther 1909S59 |--C. sordidusND13 |--C. strongylocephalus (Bleeker 1854)S59 (see below for synonymy) `--C. ultramarinus (Jordan & Seale 1906) [=Callyodon ultramarinus]S59 Labridae incertae sedis: Pseudodax Bleeker 1804 [Pseudodacinae]S57 |--*P. moluccanus (Valenciennes 1839)S57 [=Odax moluccanusM58, P. moluccensis (l. c.)JR10] |--‘Odax’ borbonicus Val. 1839S57 `--‘Odax’ pullus Val. 1839S57 Conniella apterygia Allen 1983MM09 Xenojulis margaritaceus (Macleay 1884) [=Platyglossus margaritaceus]M58 Paracheilinus flavianalis Kuiter & Allen 1999MM09 Wetmorella Fowler 1928S57 |--*W. philippina Fowler 1928 [incl. W. philippina bifasciata]S57 |--W. albofasciata Schultz & Marshall 1954S57 `--W. ocellata Schultz & Marshall 1954S57 Cirrhilabrichthys Klausewitz 1976K76 |--*C. filamentosus Klausewitz 1976K76 `--C. rubriventralis [=Cirrhilabrus rubriventralis]K76 Hologymnosus Lacepede 1802S57 `--*H. semidiscus (Lacépède 1802)S57 (see below for synonymy) Macropharyngodon Bleeker 1861S57 |--*M. geoffroy Quoy & Gaimard 1824S57 [=Platyglossus geoffroyiM58] |--M. bipartitus Smith 1957S57 |--M. meleagris (Cuvier 1839)S57 [=Julis meleagrisM58] |--M. negrosensis Herre 1932MM09 |--M. ornatus Randall 1978MM09 |--M. pardalis (Kner 1867) [=Leptojulis pardalis; incl. Platyglossus nigromaculatus Gunther 1871]S57 `--M. varialvus Smith 1957S57 Hemipteronotus Lecepede 1802S57 |--*H. quinquemaculatus Lacepede 1802S57 |--H. aneitensis (Günther 1862)S57 [=Novacula aneitensisM58] |--H. melanopus (Bleeker 1857) [incl. H. spilonotus Bleeker 1857]S57 |--H. pavoB96 |--H. pentadactylus (Linnaeus 1758)S57 (see below for synonymy) `--H. twistii (Bleeker 1856) [=Novacula twistii]M58 ChoerodonS57 CheilininaeR13 |--CheilinusS57 |--Pseudocheilinus Bleeker 1861R13, S57 | |--*P. hexataenia (Bleeker 1857) [=Cheilinus hexataenia]S57 | |--P. evanidus Jordan & Evermann 1903S57 | |--P. margaretae Smith 1955S57 | `--P. octotaenia Jenkins 1901MM09 `--Cirrhilabrus Temminck & Schlegel 1845R13, K76 |--*C. temmincki Bleeker 1852S57 |--C. cyanopleura (Bleeker 1851)PC01 |--C. exquisitus Smith 1957S57 |--C. randalli Allen 1995MM09 `--C. solorensis Bleeker 1853M58 Bolbometopon Smith 1956S56 `--*B. muricatus (Valenciennes 1839)S56 (see below for synonymy) XanothonS59 Hipposcarus Smith 1956S59 |--*H. harid (Forskål 1775)S59 (see below for synonymy) | |--H. h. haridS59 | `--H. h. vexillus Smith 1959S59 `--H. schultzi Smith 1959S59 Peaolopesia Smith 1949S57 `--P. matthaei (Valenciennes 1839) (see below for synonymy)S57 Cheiliopsis Steindachner 1863S57 `--*C. bivittatus Steindachner 1863S57 LepidaploisS57 Anchichoerops Barnard 1927S57 `--*A. natalensis (Gilchrist & Thompson 1908) [=Choerops natalensis]S57 Fowlerella Smith 1957S57 `--*F. bicolor (Fowler & Bean 1928) [=Labroides bicolor]S57 Pseudojulis Bleeker 1861S57 |--*P. girardi (Bleeker 1858) [=Julis girardi]S57 `--P. argyreogaster Gunther 1866S57 Suezia Smith 1957S57 `--*S. caudovittatus (Steindachner 1898) [=Labrichthys caudovittatus]S57 Pseudolabrus Bleeker 1862S57 `--*P. rubiginosus (Schlegel 1845) [=Labrus rubiginosus]S57 Pseudanampses Bleeker 1862S57 `--*P. geographicus (Valenciennes 1839) (see below for synonymy)S57 CossyphusS57 |--C. maxillosusS57 |--C. rufus Linn 1758S57 `--C. terophorusS57 Ampheces pterophthalmusJR10 Coricus rostratusPP64 Eocoris bloti Bannikov & Sorbini 1991P93 Phyllopharyngodon longipinnis Bellwood 1991P93 Eolabroides szjnochae (de Zigno 1887)P93 Gillidia antiqua (Agassiz 1835)P93 SymphodusP93 |--S. eocenicus (White 1926)P93 `--S. mediterraneusBM03 Malacopterus [Malacopterinae]R13 `--M. reticulatusR13 Austrolabrus maculatus (Macleay 1881)H90 NotolabrusH90 |--N. celidotusHS01 `--N. parilus (Richardson 1850)H90 Pseudojuloides elongatus Ayling & Russell 1977H90 Suezichthys cyanolaemus Russell 1985H90
*Bolbometopon muricatus (Valenciennes 1839)S59 [=Scarus muricatusS59, Callyodon muricatusS57, Pseudoscarus muricatusM58; incl. Scarus gibbosus Kuhl & van Hasselt in Valenciennes 1839 (n. n.)S56]
Cetoscarus bicolor (Rüppell 1828)S59 [=Scarus bicolorM58, Callyodon bicolorM58; incl. Pseudoscarus nigripinnis Gunther 1866S59, Scarus ophthalmistius Herre 1933S56]
*Cetoscarus pulchellus (Rüppell 1835)S59 [=Scarus pulchellusS59, Callyodon pulchellusM58, Pseudoscarus pulchellusM58]
Chlorurus microrhinos (Bleeker 1854)S59 [=Callyodon microrhinosM58, *Pseudoscarus microrhinusS56, Scarus microrhinosS59]
Chlorurus strongylocephalus (Bleeker 1854)S59 [=Scarus strongylocephalusS59, Callyodon strongylocephalusM58, Pseudoscarus strongylocephalusM58; incl. Scarus microcheilus Bleeker 1861S59, Chlorurus microcheilosS59]
Cryptotomus spinidens (Quoy & Gaimard 1824) [=Scarus spinidens; incl. Ca. brachysoma Macleay 1883, Leptoscarus brachysoma, Ca. carolinus Bleeker 1878, Ca. genistriatus Günther 1909, Ca. moluccensis Macleay 1884, Ca. waigiensis Cuvier & Valenciennes 1839]M58
Hemipteronotus pentadactylus (Linnaeus 1758)S57 [=Coryphaena pentadactylaM58, Novacula pentadactylaM58; incl. Hemipteronotus bimaculata Rupp. 1828S57, H. immaculatus Val. 1839S57, Novacula punctulatus Val. 1839M58, S57, H. temporalis Regan 1905S57, H. tessellatus Val. 1839S57]
*Hipposcarus harid (Forskål 1775)S59 [=Scarus haridS59, Callyodon haridM58; incl. C. latus Val. 1839S56, Scarus longiceps Cuvier & Valenciennes 1839M58, S. mastax Ruppell 1828S59, Callyodon ruppellii Val. 1839S56]
*Hologymnosus semidiscus (Lacépède 1802)S57 [=Labrus semidiscusS57; incl. Coris annulata Lacepede 1802M58, S57, H. annulatusH90, H. doliatus Valenciennes 1839S57, H. fasciatus Lacepede 1802S57, H. rosea Q & G 1835S57, H. semipartitus Bleeker 1875S57]
Peaolopesia matthaei (Valenciennes 1839) [=Julis matthaei; incl. Xiphocheilus gymnogenys Gunther 1866, *Peaolopesia gymnogenys]S57
*Pseudanampses geographicus (Valenciennes 1839) [=Anampses geographicus; incl. Anampsodax lienardi Bleeker & Pollen 1874, Pseudanampses lineolatus Bennett 1831]S57
*Type species of generic name indicated
References
[BM03] Barreiros, J. P., T. Morato, R. S. Santos & A. E. de Borba. 2003. Interannual changes in the diet of the almaco jack, Seriola rivoliana (Perciformes: Carangidae) from the Azores. Cybium 27 (1): 37–40.
[B96] Bond, C. E. 1996. Biology of Fishes 2nd ed. Saunders College Publishing: Fort Worth.
[B79] Bosence, D. W. J. 1979. The factors leading to aggregation and reef formation in Serpula vermicularis L. In: Larwood, G., & B. R. Rosen (eds) Biology and Systematics of Colonial Organisms pp. 299–318. Academic Press: London.
[CH97] Castro, P., & M. E. Huber. 1997. Marine Biology 2nd ed. WCB McGraw-Hill: Boston.
[D56] Dawes, B. 1956. The Trematoda with special reference to British and other European forms. University Press: Cambridge.
[F15] Fowler, H. W. 1915. Cold-blooded vertebrates from Florida, the West Indies, Costa Rica, and eastern Brazil. Proceedings of the Academy of Natural Sciences of Philadelphia 67 (2): 244–269.
[F16] Fowler, H. W. 1916. The fishes of Trinidad, Grenada, and St. Lucia, British West Indies. Proceedings of the Academy of Natural Sciences of Philadelphia 67 (3): 520–548.
[G75] Grant, E. M. 1975. Guide to Fishes. The Co-ordinator-General’s Department: Brisbane (Australia).
[HS01] Hayward, B. W., A. B. Stephenson, M. S. Morley, W. M. Blom, H. R. Grenfell, F. J. Brook, J. L. Riley, F. Thompson & J. J. Hayward. 2001. Marine biota of Parengarenga Harbour, Northland, New Zealand. Records of the Auckland Museum 37: 45–80.
[H90] Hutchins, J. B. 1990. Fish survey of South Passage, Shark Bay, Western Australia. In: Berry, P. F., S. D. Bradshaw & B. R. Wilson (eds) Research in Shark Bay: Report of the France-Australe Bicentenary Expedition Committee pp. 263–278. Western Australian Museum.
[JR10] Jordan, D. S., & R. E. Richardson. 1910. Check-list of the species of fishes known from the Philippine archipelago. Bureau of Printing: Manila.
[K76] Klausewitz, W. 1976. Cirrhilabrichthys filamentosus n. gen., n. sp., aus der Javasee (Pisces: Labridae). Senckenbergiana Biologica 57 (1–3): 11–14.
[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.
[MM09] Moore, G., & S. Morrison. 2009. Fishes of three North West Shelf atolls of Western Australia: Mermaid (Rowley Shoals), Scott and Seringapatam Reefs. Records of the Western Australian Museum Supplement 77: 221–255.
[M58] Munro, I. S. R. 1958. The fishes of the New Guinea region: a check-list of the fishes of New Guinea incorporating records of species collected by the Fisheries Survey Vessel “Fairwind” during the years 1948 to 1950. Papua and New Guinea Agricultural Journal 10 (4): 97–369 (reprinted: 1958. Territory of Papua and New Guinea Fisheries Bulletin no. 1).
[M02] Munroe, T. A. 2002. Wrasses. Family Labridae. In: Collette, B. B., & G. Klein-MacPhee (eds) Bigelow and Schroeder’s Fishes of the Gulf of Maine 3rd ed. pp. 448–466. Smithsonian Institute Press: Washington.
[ND13] Near, T. J., A. Dornburg, R. I. Eytan, B. P. Keck, W. L. Smith, K. L. Kuhn, J. A. Moore, S. A. Price, F. T. Burbrink, M. Friedman & P. C. Wainwright. 2013. Phylogeny and tempo of diversification in the superradiation of spiny-rayed fishes. Proceedings of the National Academy of Sciences of the USA 110 (31): 12738–12743.
[P93] Patterson, C. 1993. Osteichthyes: Teleostei. In: Benton, M. J. (ed.) The Fossil Record 2 pp. 621–656. Chapman & Hall: London.
[PP64] Peres, J. M., & J. Picard. 1964. Nouveau manuel de bionomie benthique de la mer Mediterranee. Recueil des Travaux de la Station Marine d’Endoume, Bulletin 31 (27): 5–137.
[PC01] Pichelin, S., & T. H. Cribb. 2001. The status of the Diplosentidae (Acanthocephala: Palaeacanthocephala) and a new family of acanthocephalans from Australian wrasses (Pisces: Labridae). Folia Parasitologica 48: 289–303.
[R13] Regan, C. T. 1913. The classification of the percoid fishes. Annals and Magazine of Natural History, series 8, 12: 111–145.
[S56] Smith, J. L. B. 1956. The parrot fishes of the family Callyodontidae of the western Indian Ocean. Ichthyological Bulletin 1: 1–23 (reprinted: 1969, with notes).
[S57] Smith, J. L. B. 1957. List of fishes of the family Labridae in the western Indian Ocean. Ichthyological Bulletin 7: 99–114.
[S59] Smith, J. L. B. 1959. The identity of Scarus gibbus Ruppell, 1828 and of other parrotfishes of the family Callyodontidae from the Red Sea and the western Indian Ocean. Ichthyological Bulletin 16: 265–282.