Belongs within: Leiodinae.
The Leiodini are a group of round fungus beetles characterised by a 5-5-4 tarsal formula (Peck 2001).
Characters (from Peck 2001): Head without occipital carina; labrum more or less deeply emarginate apically; cervical sclerites present; prosternum in front of coxae much shorter than coxal width; hind coxae contiguous; tarsal formula 5-5-4.
LeiodiniP90
|--Anogdus LeConte 1866 [incl. Neocyrtusa Brown 1937]P01
|--Liocyrtusa Daffner 1982P01
|--Ecarinosphaerula Hatch 1929P90
| `--E. ecarina Hatch 1929P01
|--Lionothus Brown 1937 [incl. Pseudocyrtusa Portevin 1942]P01
| `--L. ulkeiP90
|--Zeadolopus Broun 1903 [incl. Apheloplastus Brown 1937]P01
| `--Z. egenus [=Apheloplastus egenus]P90
|--Isoplastus Horn 1880P90
| `--I. fossor Horn 1880P01
|--Leiodes Latreille 1796P01 (see below for synonymy)
| |--L. assimilisP01
| |--L. ferrugineusG20 [=Anisotoma ferrugineaL02, Diaperis (Leiodes) ferrugineaG20]
| `--L. paludicolaLC20
`--Cyrtusa Erichson 1842 [incl. Caenocyrta Brown 1937]P01
|--C. luggeriP90
`--‘Caenocyrta’ picipennisP90
Leiodes Latreille 1796 [=Liodes Erichson 1845 non Heyden 1826; incl. Eremosphaerula Hlisnikovsky 1967, Oosphaerula Ganglbauer 1896, Oreosphaerula Ganglbauer 1899, Parahydnobius Fleischer 1908, Pseudohydnobius Ganglbauer 1899, Pteromerula Fleischer 1905, Trichosphaerula Fleischer 1905]P01
*Type species of generic name indicated
References
[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.
[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.
[LC20] Lü, L., C.-Y. Cai, X. Zhang, A. F. Newton, M. K. Thayer & H.-Z. Zhou. 2020. Linking evolutionary mode to palaeoclimate change reveals rapid radiations of staphylinoid beetles in low-energy conditions. Current Zoology 66 (4): 435–444.
[P90] Peck, S. B. 1990. Insecta: Coleoptera Silphidae and the associated families Agyrtidae and Leiodidae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1113–1136. John Wiley & Sones: New York.
[P01] Peck, S. B. 2001. Leiodidae Fleming, 1821. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 250–258. CRC Press: Boca Raton.