Leucosiidae

Ebalia tumefacta, copyright B. E. Picton.

Belongs within: Heterotremata.

The Leucosiidae, purse or pebble crabs, are a group of crabs with a rounded or pentagonal, usually very convex, carapace. The majority of species are found on coastal sand bottoms, gravels and mud.

<==Leucosiidae
    |--Heteronucia venusta Nobili 1906TSH09
    |--Myrine kessleri (Paul’son 1875)MG-H11
    |--Parilia major Sakai 1962MG-H11
    |--Philyra pisumMG-H11, CG97
    |--Typilobus trispinosus Lörenthey 1909BWW93
    |--Ixoides cornutus MacGilchrist 1905J90
    |--Pseudophilyra perryi (Miers 1877) [=Leucosia perryi]J90
    |--Merocryptus Milne Edwards 1873B64
    |    `--M. lambriformis Milne Edwards 1873 [incl. Ebalia rugulosa Yokoya 1933]B64
    |--OreophorusMG-H11
    |    |--O. morum (Alcock 1896)J90
    |    `--O. reticulatus Adams & White 1849MG-H11
    |--RandalliaMG-H11
    |    |--R. eburnea Alcock 1896MG-H11
    |    `--R. pustuloides Sakai 1961MG-H11
    |--IxaJ90
    |    |--I. acuta Tyndale-Biscoe & George 1962J90
    |    `--I. monodi Holthuis & Gottlieb 1956KK03
    |--Persephona Leach 1817BF07
    |    |--P. lichtensteinii Leach 1817BF07
    |    |--P. mediterranea (Herbst 1794)BF07
    |    `--P. punctata (Linnaeus 1758)BF07
    |--MyraMG-H11
    |    |--M. affinis Bell 1855J90
    |    |--M. australis Haswell 1880J90
    |    |--M. curtimana Galil 2001MG-H11
    |    |--M. fugaxP71
    |    |--M. mammillaris Bell 1855J90
    |    `--M. subgranulata (Fabricius 1798)KK03
    |--ArcaniaMG-H11
    |    |--A. cornuta (MacGilchrist 1905)MG-H11
    |    |--A. elongata Yokoya 1933MG-H11
    |    |--A. gracilis (Henderson 1898)MG-H11
    |    |--A. muricata Galil 2002MG-H11
    |    |--A. novemspinosa (Adams & White 1849)MG-H11
    |    |--A. quinquespinosa Alcock & Anderson 1894J90
    |    |--A. septemspinosa (Fabricius 1787)MG-H11
    |    `--A. undecimspinosa de Haan 1841J90
    |--Ebalia Leach 1817B64
    |    |--E. algirica Lucas 1846E12
    |    |--E. cranchii Leach 1817KK03
    |    |--E. deshayesii Lucas 1846E12
    |    |--E. edwardsii Costa 1838KK03
    |    |--E. granulosa Milne-Edwards 1837KK03
    |    |--E. insignis Lucas 1846E12
    |    |--E. laevis (Bell 1855) [=Phlyxia laevis; incl. P. cheesmani Filhol 1886, E. cheesmani]B64
    |    |--E. nux Milne-Edwards 1883KK03
    |    |--E. tuberculosa (Milne Edwards 1873) [=Persephona tuberculosa; incl. Phlyxia granulosa Haswell 1880]B64
    |    |--E. tuberosa (Pennant 1777)KK03
    |    `--E. tumefactaM62
    `--LeucosiaL02
         |--L. craniolarisL02
         |--L. haematosticta Adams & White 1848MG-H11
         |--L. haswelli Miers 1886J90
         |--L. nucleusG20
         |--L. ocellata Bell 1855MG-H11
         |--L. perlata de Haan 1841J90
         |--L. pubescens Miers 1877J90
         |--L. reticulata Miers 1877J90
         |--L. scabriusculaR26
         |--L. signata Paulson 1975KK03
         |--L. unidentata de Haan 1841J90
         `--L. whitei Bell 1855MG-H11

*Type species of generic name indicated

References

[B64] Bennett, E. W. 1964. The marine fauna of New Zealand: Crustacea Brachyura. New Zealand Department of Scientific and Industrial Research Bulletin 153: 1–120.

[BF07] Braga, A. A., A. Fransozo, G. Bertini & P. B. Fumis. 2007. Bathymetric distribution and recruitment of the spider crab Libinia spinosa H. Milne Edwards 1834 in the Ubatuba and Caraguatatuba regions, northern coast of São Paulo, Brazil (Crustacea, Brachyura, Majoidea, Pisidae). Senckenbergiana Biologica 87 (1): 7–16.

[BWW93] Briggs, D. E. G., M. J. Weedon & M. A. Whyte. 1993. Arthropoda (Crustacea excluding Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 321–342. Chapman & Hall: London.

[CG97] Cohen, B. L., & A. B. Gawthrop. 1997. The brachiopod genome. In: Kaesler, R. L. (ed.) Treatise on Invertebrate Paleontology pt H. Brachiopoda, Revised vol. 1. Introduction pp. 189–211. The Geological Society of America: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[J90] Jones, D. S. 1990. Annotated checklist of marine decapod Crustacea from Shark Bay, Western Australia. In: Berry, P. F., S. D. Bradshaw & B. R. Wilson (eds) Research in Shark Bay: Report of the France-Australe Bicentenary Expedition Committee pp. 169–208. Western Australian Museum.

[KK03] Kocataş, A., & T. Katağan. 2003. The decapod crustacean fauna of the Turkish seas. Zoology in the Middle East 29: 63–74.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[M62] Monniot, F. 1962. Recherches sur les graviers a Amphioxus de la région de Banyuls-sur-Mer. Vie et Milieu 13: 231–322.

[P71] Por, F. D. 1971. One hundred years of Suez Canal—a century of Lessepsian migration: retrospect and viewpoints. Systematic Zoology 20 (2): 138–159.

[R26] Risso, A. 1826. Histoire naturelle des principales productions de l’Europe méridionale et particulièrement de celles des environs de Nice et des Alpes maritimes vol. 5. F.-G. Levrault: Paris.

[TSH09] Titelius, M. A., A. Sampey & C. G. Hass. 2009. Crustaceans of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, north Western Australia. Records of the Western Australian Museum Supplement 77: 145–176.

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