Liacarus (Liacarus)

Liacarus subterraneus, copyright R. Penttinen.

Belongs within: Gustavioidea.

The type subgenus of the genus Liacarus is a subcosmopolitan group of oribatid mites found in soil litter. The type species, L. nitens, is a member of a group of species with narrowly fused lamellae bearing narrow cusps and a sharply pointed interlamellar mucro.

Characters (from Balogh & Balogh 1992): Prodorsum with narrow lamellae, converging apically, lateral parts of prodorsum usually uncovered; sensillus fusiform. Humeral region of notogaster with two pairs of setae c1 and c2 near each other. Genital plates with six pairs of setae. Legs tridactyle.

<==Liacarus Michael 1898 (Liacarus) (see below for synonymy)S04
    |--*L. (L.) nitens (Gervais 1844)S04 [=Oribata nitensS04, *Leiosoma nitensCH98]
    |--L. (L.) arduus Fujikawa 1989S04
    |--L. (L.) atraktus (Woolley & Higgins 1966)S04 [=*Stenoxenillus atraktusBB92, Stonyxenillus atraktusW72]
    |--L. (L.) bidentatus Ewing 1918S04
    |--L. (L.) borealis Wen 1991S04
    |--L. (L.) brevilamellatus Mihelčič 1955 (see below for synonymy)S04
    |--L. (L.) ‘castaneus’ (Banks 1906) [=Notaspis castanea non Hermann 1804]S04
    |--L. (L.) celisi Balogh 1958 (n. d.)S04
    |--L. (L.) chiebunensis Fujita & Fujikawa 1984S04
    |--L. (L.) chroniosus Woolley 1968 [incl. L. robustus Mihelčič 1954 non Ewing 1918]S04
    |--L. (L.) cidarus Woolley 1968S04
    |--L. (L.) conjunctus Mihelčič 1954S04
    |--L. (L.) contiguus Aoki 1969S04
    |--L. (L.) coracinus (Koch 1841)S04 [=Oribata coracinaS04, Leiosoma coracinumBA85]
    |    |--L. c. coracinusS04 (see below for synonymy)
    |    `--L. c. simplex Mihelčič 1952S04
    |--L. (L.) curvidentatus Balogh 1958 (n. d.)S04
    |--L. (L.) detosus Woolley 1968S04
    |--L. (L.) emeiensis Wen 1991S04
    |--L. (L.) externus Aoki 1987S04
    |--L. (L.) flammeus Aoki 1967S04
    |--L. (L.) gammatus Aoki 1967S04
    |    |--L. g. gammatusS04
    |    `--L. g. coreanus Choi 1994S04
    |--L. (L.) granulatus Willmann 1940 (n. d.)S04
    |--L. (L.) inermis Aoki 1965S04
    |--L. (L.) janetscheki Mihelčič 1957S04
    |--L. (L.) jordanoi Moraza & Pérez-Íñigo 1979S04
    |--L. (L.) kanekoi Bayartogtokh & Aoki 2002S04
    |--L. (L.) kilchini Choi 1985S04
    |--L. (L.) koeszegiensis Balogh 1943 [incl. L. sejunctus Mihelčič 1954]S04
    |--L. (L.) laterostris Mihelčič 1954S04
    |--L. (L.) latilamellatus Kaneko & Aoki 1982S04
    |--L. (L.) latus Ewing 1909S04
    |--L. (L.) lectronus Higgins & Woolley 1969S04
    |--L. (L.) leleupi Balogh 1958 (n. d.)S04
    |--L. (L.) lencoranicus Krivolutsky 1967S04
    |--L. (L.) logipilis (Moniez 1894) [=Leiosoma logipilis]S04
    |--L. (L.) longilamellatus Mihelčič 1954S04
    |--L. (L.) lucidus Ewing 1909S04
    |--L. (L.) luscus Hirauchi 1998S04
    |--L. (L.) madeirensis Willmann 1939S04
    |--L. (L.) marginatus (Nicolet 1855) [=Leiosoma marginatum]S04
    |--L. (L.) matshabelii Djaparidze 1980S04
    |--L. (L.) medialis Banks 1910S04
    |--L. (L.) montanus Aoki 1984S04
    |--L. (L.) mucronatus Willmann 1939S04
    |--L. (L.) murotensis Aoki 1988S04
    |--L. (L.) nigrescens Pearce 1906S04
    |--L. (L.) nitidulus Krivolutsky 1967S04
    |--L. (L.) ocellatus Aoki 1987S04
    |--L. (L.) olivaceus (Grube 1859) (n. d.) [=Leiosoma olivaceum]S04
    |--L. (L.) oribatelloides Winkler 1956S04
    |--L. (L.) orthogonios Aoki 1959S04
    |--L. (L.) ovatus Mihelčič 1954 (see below for synonymy)S04
    |--L. (L.) parvus Mihelčič 1954S04
    |--L. (L.) perezinigoi Capilla 1972S04
    |--L. (L.) pseudocontiguus Fujikawa 1991S04
    |--L. (L.) regeli Djaparidze 1983S04
    |--L. (L.) reticulatus Mahunka 1966S04
    |--L. (L.) robustus Ewing 1918S04
    |--L. (L.) rotundatus Mihelčič 1954S04
    |--L. (L.) schweizeri Forsslund 1963 [incl. L. latus Schweizer 1956 non Ewing 1909]S04
    |--L. (L.) sphaericus Djaparidze 1980S04
    |--L. (L.) spiniger Jacot 1937S04
    |--L. (L.) subterraneus (Koch 1844)S04 (see below for synonymy)
    |--L. (L.) tanzicus Mahunka 1983S04
    |--L. (L.) tremellae (Linnaeus 1761) (n. d.)S04 (see below for synonymy)
    |--L. (L.) trichionus (Woolley & Higgins 1966) [=Leuroxenillus trichionus]S04
    |--L. (L.) tubifer Djaparidze & Melamud 1990S04
    |--L. (L.) unjangensis Choi, Bayartogtokh & Aoki 2000S04
    |--L. (L.) willmanni Pschorn-Walcher 1951S04
    |--L. (L.) xylariae (Schrank 1803) (see below for synonymy)S04
    `--L. (L.) yayeyamensis Aoki 1973S04

Liacarus Michael 1898 (Liacarus) [=Leiosoma Nicolet 1855 nec Stephens 1829 nec Chevrolat 1837; incl. Leuroxenillus Woolley & Higgins 1966, Stenoxenillus Woolley & Higgins 1966]S04

Liacarus (Liacarus) brevilamellatus Mihelčič 1955 [incl. L. cartalinicus Djaparidze 1979, L. internodentatus Kulijev 1962, L. major Mihelčič 1955, L. ovalis Mihelčič 1956, L. vastus Mihelčič 1956]S04

Liacarus (Liacarus) coracinus coracinus (Koch 1841)S04 [incl. Oribata politus Koch & Berendt 1854S04, Leiosoma similis Nicolet 1855S04, Liacarus similisG41, Li. vombi Dalenius 1950S04]

Liacarus (Liacarus) ovatus Mihelčič 1954 non Oribata ovata Koch 1839 (not preoc. if in dif. gen.) [incl. L. mihelcici Woolley 1968]S04

Liacarus (Liacarus) subterraneus (Koch 1844)S04 [=Oribata subterraneaS04; incl. Li. gracilis Mihelčič 1954S04, Leiosoma lativentris Nicolet 1855S04, Li. lativentrisM98]

Liacarus (Liacarus) tremellae (Linnaeus 1761) (n. d.)S04 [=Acarus tremellaeS04; incl. Oribata globosa Koch 1844S04, Conoppia globosusG36, Leiosoma globosumM98, Liacarus globosusG36]

Liacarus (Liacarus) xylariae (Schrank 1803) [=Acarus xylariae; incl. L. cuspidatus Mihelčič 1954, Oribata fusca Koch 1841, L. translamellatus Kulijev 1962]S04

*Type species of generic name indicated

References

[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[BA85] Bernini, F., & R. Arcidiacono. 1985. Notulae oribatologicae XXXVI. A catalogue of the Sicilian oribatids (Acarida, Oribatida). Animalia 12 (1–3): 165–205.

[CH98] Colloff, M. J.. & R. B. Halliday. 1998. Oribatid Mites: A catalogue of Australian genera and species. CSIRO Publishing: Collingwood.

[G36] Grandjean, F. 1936. Observations sur les oribates (10e série). Bulletin du Muséum, 2e série, 8 (3): 246–253.

[G41] Grandjean, F. 1941. Statistique sexuelle et parthénogenèse chez les oribates (acariens). Comptes Rendus des Séances de l’Académie des Sciences 212: 463–466.

[M98] Michael, A. D. 1898. Oribatidae. In: H. Lohmann (ed.) Das Tierreich. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen vol. 3. Acarina pp. 1–93. R. Friedländer und Sohn: Berlin.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

[W72] Woolley, T. A. 1972. The systematics of the Liacaroidea (Acari: Cryptostigmata). Acarologia 14 (2): 250–257.

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