Linyphiidae

Porrhomma pygmaeum, from Pavouci.

Belongs within: Araneoidea.
Contains: Lepthyphantes, Mughiphantes, Agyneta, Haplinis, Parafroneta, Diplocephalus, Erigone, WalckenaeriaOedothorax, Linyphia, Laetesia, Porrhomma, Centromerus, Bathyphantes.

The Linyphiidae are a hyperdiverse family of small spiders, most of which build delicate sheet webs among vegetation. They include the Erigoninae, money spiders (sometimes treated as a separate family), very small spiders that are usually dark and shiny, and often have the frontal region of males modified with lobes, pits or setal fields (Jocqué & Dippenaar-Schoeman 2007). The Linyphiinae are united by the presence in males of the lamella characteristica, a sclerite of the embolic division of the pedipalp. Members of the Mynogleninae have characteristic cephalic pits just below the anterior lateral eyes, as well as numerous trichobothria on the metatarsi. The Micronetinae have a complex embolic division on the male pedipalp, often with the sperm duct dilated at the base of the embolus.

Characters (from Jocqué & Dippenaar-Schoeman 2007): Very small to small; eight eyes present; labium rebordered; chelicerae usually with stridulating file; numerous teeth, mostly three or four, on fang furrow; legs slender, provided with setae, especially on tibiae and metatarsi; autospasy at patella-tibia joint; tarsi cylindrical, with three tarsal claws; tarsus IV without comb; ecribellate; entelegyne; male palp with often U-shaped intersegmental paracymbium.

<==Linyphiidae
    |--MicronetinaeTS06
    |    |--LepthyphantesTS06
    |    |--MughiphantesTS06
    |    |--AgynetaTS06
    |    |--Microneta Menge 1869W76
    |    |    `--M. varia (Blackwall 1841)MKD01 [incl. M. nicholsonii Pickard-Cambridge 1906V09, P-C06]
    |    |--Ascetophantes Tanasevitch & Saaristo 2006TS06
    |    |    `--*A. asceticus (Tanasevitch 1987) [=Lepthyphantes asceticus]TS06
    |    |--Fistulaphantes Tanasevitch & Saaristo 2006TS06
    |    |    `--*F. canalis Tanasevitch & Saaristo 2006TS06
    |    |--Spiralophantes Tanasevitch & Saaristo 2006TS06
    |    |    `--*S. mirabilis Tanasevitch & Saaristo 2006TS06
    |    |--Piniphantes Saaristo & Tanasevitch 1996TS06
    |    |    `--P. himalayensis (Tanasevitch 1987) [=Lepthyphantes uzbekistanicus himalayensis]TS06
    |    |--Birgerius Saaristo 1973MKD01
    |    |    `--*B. microps (Simon 1911) [=Centromerus microps]MKD01
    |    |--Claviphantes Tanasevitch & Saaristo 2006TS06
    |    |    |--*C. bifurcatus (Tanasevitch 1987) [=Lepthyphantes bifurcatus]TS06
    |    |    `--C. bifurcatoides (Tanasevitch 1987) [=Lepthyphantes bifurcatoides]TS06
    |    |--Megalepthyphantes Wunderlich 1994TS06
    |    |    |--M. nebulosoides (Wunderlich 1977) [=Lepthyphantes nebulosoides]TS06
    |    |    `--M. nebulosus (Sundevall 1830)TS06
    |    |--Martensinus Wunderlich 1973TS06
    |    |    |--M. annulatus Wunderlich 1973TS06
    |    |    `--M. micronetiformis Wunderlich 1973TS06
    |    |--Indophantes Saaristo & Tanasevitch 2003TS06
    |    |    |--I. agamus Tanasevitch & Saaristo 2006TS06
    |    |    |--I. bengalensis Saaristo & Tanasevitch 2003TS06
    |    |    `--I. digitulus (Thaler 1987) [=Lepthyphantes digitulus]TS06
    |    |--Anguliphantes Saaristo & Tanasevitch 1996TS06
    |    |    |--A. dybowskii (P.-Cambridge 1873)MKD01
    |    |    |--A. karpinskii (P.-Cambridge 1873)MKD01
    |    |    `--A. nepalensis (Tanasevitch 1987) [=Lepthyphantes nepalensis]TS06
    |    |--Palliduphantes Saaristo & Tanasevitch 2001TS06
    |    |    |--P. ericaeus (Blackwall 1853)BBM02
    |    |    |--P. pallidus (Cambridge 1871)BBM02
    |    |    `--P. theosophicus (Tanasevitch 1987) [=Lepthyphantes theosophicus]TS06
    |    |--Himalaphantes Tanasevitch 1992TS06
    |    |    |--H. grandiculus (Tanasevitch 1987) [=Lepthyphantes grandiculus]TS06
    |    |    |--H. magnus (Tanasevitch 1987) [=Lepthyphantes magnus]TS06
    |    |    `--H. martensi (Thaler 1987) [=Lepthyphantes martensi]TS06
    |    `--Tenuiphantes Saaristo & Tanasevitch 1996TS06
    |         |--T. alacris (Blackwall 1853)MKD01
    |         |--T. altimontanus Tanasevitch & Saaristo 2006TS06
    |         |--T. crassus Tanasevitch & Saaristo 2006TS06
    |         |--T. mengei (Kulczyński 1887)MKD01
    |         |--T. plumipes (Tanasevitch 1987) [=Lepthyphantes plumipes]TS06
    |         |--T. tenuis (Blackwall 1852)PVD10
    |         `--T. zimmermanni (Betkau 1890)BBM02
    |--MynogleninaePVD10
    |    |--HaplinisPVD10
    |    |--ParafronetaPVD10
    |    |--Cassafroneta Blest 1979PVD10
    |    |    `--*C. forsteri Blest 1979PVD10
    |    |--Hyperafroneta Blest 1979PVD10
    |    |    `--*H. obscura Blest 1979PVD10
    |    |--Poecilafroneta Blest 1979PVD10
    |    |    `--*P. caudata Blest 1979PVD10
    |    |--Protoerigone Blest 1979PVD10
    |    |    |--*P. otagoa Blest 1979PVD10
    |    |    `--P. obtusa Blest 1979PVD10
    |    |--Megafroneta Blest 1979PVD10
    |    |    |--*M. elongata Blest 1979PVD10
    |    |    |--M. dugdaleae Blest & Vink 2002PVD10
    |    |    `--M. gigas Blest 1979PVD10
    |    |--Metafroneta Blest 1979PVD10
    |    |    |--*M. sinuosa Blest 1979PVD10
    |    |    |--M. minima Blest 1979PVD10
    |    |    `--M. subversa Blest & Vink 2002PVD10
    |    |--Mynoglenes Simon 1905 [incl. Afroneta Holm 1968, Paralinyphia Bryant 1933]W76
    |    |    |--M. dilorisM83
    |    |    |--M. insolens Simon 1905F64 [incl. Linyphia antipodiana Berland 1925H39]
    |    |    |--M. majorFF99
    |    |    |--M. marrineri Hogg 1909W76
    |    |    |--M. simplicipalpis Wunderlich 1976W76
    |    |    `--M. titanFF99
    |    |--Novafroneta Blest 1979PVD10
    |    |    |--*N. vulgaris Blest 1979PVD10
    |    |    |--N. annulipes Blest 1979PVD10
    |    |    |--N. gladiatrix Blest 1979PVD10
    |    |    |--N. nova Blest & Vink 2003PVD10
    |    |    |--N. parmulata Blest 1979PVD10
    |    |    `--N. truncata Blest & Vink 2003PVD10
    |    |--Promynoglenes Blest 1979PVD10
    |    |    |--*P. nobilis Blest 1979PVD10
    |    |    |--P. grandis Blest 1979PVD10
    |    |    |--P. minuscula Blest & Vink 2003PVD10
    |    |    |--P. minuta Blest & Vink 2002PVD10
    |    |    |--P. parvula Blest 1979PVD10
    |    |    `--P. silvestris Blest 1979PVD10
    |    |--Pseudafroneta Blest 1979PVD10
    |    |    |--*P. incerta (Bryant 1935)PVD10
    |    |    |--P. frigida Blest 1979PVD10
    |    |    |--P. lineata Blest 1979PVD10
    |    |    |--P. maxima Blest 1979NS00
    |    |    |--P. pallida Blest 1979PVD10
    |    |    |--P. perplexa Blest 1979PVD10
    |    |    `--P. prominula Blest 1979NS00
    |    `--Metamynoglenes Blest 1979PVD10
    |         |--*M. incurvata Blest 1979PVD10
    |         |--M. absurda Blest & Vink 2002PVD10
    |         |--M. attenuata Blest 1979PVD10
    |         |--M. flagellata Blest 1979PVD10
    |         |--M. gracilis Blest 1979PVD10
    |         |--M. helicoidesPVD10
    |         |--M. magna Blest 1979PVD10
    |         `--M. ngongotaha Blest & Vink 2002PVD10
    |--Erigoninae [Erigonidae]W76
    |    |--Eperigone tridentataM83
    |    |--DiplocephalusMKD01
    |    |--ErigoneMKD01
    |    |--Tunagyna debilisJVHN90
    |    |--WalckenaeriaD90
    |    |--Delorrhipis fronticornis Simon 1884MKD01
    |    |--Sciastes Bishop & Crosby 1938W76
    |    |--Aphileta Hull 1920W76
    |    |--CeratinopsD90
    |    |--EridantesD90
    |    |--Minyriolus pusillus (Wider 1834)D90, MKD01
    |    |--IslandianaD90
    |    |--Halorates [incl. Catabrithorax]D90
    |    |--SpirembolusD90
    |    |--SisicottusD90
    |    |--Diplocentria [incl. Scotoussa]D90
    |    |--ScylaceusD90
    |    |--EogonatiumP92
    |    |--OedothoraxTS06
    |    |--Microctenonyx Dahl 1886PVD10
    |    |    `--M. subitaneus (Pickard-Cambridge 1875)PVD10
    |    |--Holminaria Eskov 1991MKD01
    |    |    `--H. sibirica Eskov 1991 [incl. Birgerius triangulus Tao, Li & Zhu 1995]MKD01
    |    |--Uusitaloia Marusik, Koponen & Danilov 2001MKD01
    |    |    `--*U. transbaicalica Marusik, Koponen & Danilov 2001MKD01
    |    |--Lessertia Smith 1908PVD10
    |    |    `--*L. dentichelis (Simon 1884)PVD10
    |    |--Ceratinopsis Emert. 1882D90, S06
    |    |    |--C. aliena Kulcz. 1885S06
    |    |    `--C. interventa Chamberlin 1948D90, MKD01
    |    |--Mermessus Pickard-Cambridge 1899PVD10
    |    |    |--M. denticulatus (Banks 1898)PO08
    |    |    `--M. fradeorum (Berland 1932)PVD10
    |    |--ScotinotylusD90
    |    |    |--‘Scironis’ autorC90
    |    |    `--S. protervus (Koch 1879)MKD01
    |    |--CollinsiaD90
    |    |    |--C. inerrans (Cambridge 1855)BBM02
    |    |    `--C. submissa (Koch 1879)MKD01
    |    |--Araeoncus Simon 1884PVD10
    |    |    |--*A. humilis (Blackwall 1841)PVD10
    |    |    |--A. excelsus (Koch 1880)S06
    |    |    `--A. ruderalis Sörensen 1898S06
    |    |--GrammonotaD90
    |    |    |--G. angustaJDW90
    |    |    |--G. inornata Emerton 1882WCH11
    |    |    `--G. pictilisJDW90
    |    |--HypselistesJVHN90
    |    |    |--H. florensJVHN90
    |    |    |--H. jacksoni (P.-Cambridge 1902)MKD01
    |    |    `--H. semiflavus (Koch 1879)MKD01
    |    |--Pocadicnemis Simon 1884C90, S06
    |    |    |--P. americana Millidge 1976PS08
    |    |    |--P. juncea Locket & Millidge 1953PG08
    |    |    `--P. pumila (Blackwall 1841)MKD01
    |    |--Tapinocyba Simon 1884D90, S06
    |    |    |--T. aquilonaris (Koch 1880)S06
    |    |    |--T. becki (Cambr. 1871)S06
    |    |    |--T. insecta (Koch 1869)BBM02
    |    |    `--T. pallens (Cambr. 1871)S06
    |    |--SavigniaMKD01
    |    |    |--S. boreaMKD01
    |    |    |--S. eskovi Marusik, Koponen & Danilov 2001MKD01
    |    |    |--S. frontata (Blackwall 1833)BH02
    |    |    `--S. nenilini Marusik 1988MKD01
    |    |--Micryphantes Koch 1833P92, PL73 [Micryphantidae]
    |    |    |--*M. ovatus Koch 1833 [=Meioneta ovata]PL73
    |    |    |--M. corniger (Blackw. 1856)S06
    |    |    |--M. decora (Cambr. 1871)S06
    |    |    |--M. rurestris Koch 1836S06
    |    |    `--M. sublimis (Cambr. 1875)S06
    |    |--Ceraticelus Simon 1884D90, S06
    |    |    |--C. atricepsJDW90
    |    |    |--C. emertoni (Pickard-Cambridge 1874)MF11
    |    |    |--C. fissicepsJVHN90
    |    |    |--C. innominabilis Crosby 1905S06
    |    |    `--C. orientalis Eskov 1987MKD01
    |    `--Ceratinella Emert. 1882D90, S06
    |         |--C. brevipes (Westring 1851)MKD01
    |         |--C. brevis (Wider 1834)BBM02
    |         |--C. oculatissima Strand 1901S06
    |         |--C. rotundaK01
    |         |--C. scabrosa (P.-Cambridge 1863)BBM02
    |         `--C. sydneyensis Wunderlich 1976W76
    `--LinyphiinaeW76
         |--MioxenaC90
         |--Tennesseellum formicum (Emerton 1882)WCH11
         |--LinyphiaW76
         |--LaetesiaPVD10
         |--PorrhommaH39
         |--Dicyphus Menge 1866S06
         |--CentromerusS06
         |--BathyphantesS06
         |--Gongylidium Menge 1867S06
         |    `--G. rufipes (Linnaeus 1758)BBM02
         |--Microlinyphia Gerhardt 1928W76
         |    `--M. pusilla (Sundevall 1830)MKD01 [=Linyphia pusillaW76]
         |--Ostearius Hull 1911PVD10
         |    `--O. melanopygius (Pickard-Cambridge 1879)W76 (see below for synonymy)
         |--Dicymbium Menge 1867S06
         |    `--D. nigrum (Blackwall 1834)BBM02
         |--Pseudogonatium Strand 1901S06
         |    `--P. fuscomarginatum Strand 1901S06
         |--Notioscopus Simon 1884S06
         |    `--N. curvitarsus Sörens. 1898S06
         |--Nematogmus Simon 1884S06
         |    `--N. obscurus (Blackw. 1834)S06
         |--Semljicola Strand 1906S06
         |    `--*S. barbigera (Koch 1880) [=Erigone barbigera]S06
         |--Utopiellum Strand 1901S06
         |    `--U. mirabile (Koch 1880)S06
         |--Leptorrhoptrum Kulcz. 1894S06
         |    `--L. huthwaiti (Cambr. 1861)S06
         |--Mengea Cambr. 1903S06
         |    `--M. scopigera (Grube 1859)S06
         |--Centromeria Strand 1901S06
         |    `--C. bicolor (Blackw. 1833)S06
         |--Diplostyla Emert. 1882S06
         |    `--D. concolor (Wider 1834)MF11
         |--Hemiphantes Strand 1901S06
         |    `--H. arcticus Strand 1901S06
         |--Bolephthyphantes Strand 1901S06
         |    `--B. index (Thorell 1856)S06
         |--Tarsiphantes Strand 1903–1905S06
         |    `--T. latithorax Strand 1903–1905S06
         |--Scironis Bishop & Crosby 1938C90
         |    |--S. simaC90
         |    `--S. tarsalisC90
         |--RinginaF64
         |    |--R. antarctica (Hickman 1939) [=Drapetisca antarctica]F64
         |    `--R. crozetensis Tambs-Lyche 1954F64
         |--Dismodicus Simon 1884S06
         |    |--D. bifrons (Blackwall 1841)MKD01
         |    `--D. elevatus (Koch 1838)S06
         |--Novalaetesia Millidge 1988PVD10
         |    |--*N. anceps Millidge 1988PVD10
         |    `--N. atra Blest & Vink 2003PVD10
         |--Brachycentrum Dahl 1886S06
         |    |--B. elongatum (Wid. 1834)S06
         |    `--B. simile (Sörens. 1898)S06
         |--Erigonella Dahl 1901S06
         |    |--E. groenlandica Strand 1903–1905S06
         |    `--E. hiemalis (Blackwall 1841)BH02
         |--Tiso Simon 1884S06
         |    |--T. aestivus (Koch 1872)S06
         |    `--T. vagans (Blackwall 1834)BBM02
         |--Acartauchenius Simon 1884S06
         |    |--A. pilifrons (Koch 1880)S06
         |    `--A. scurrilis (P.-Cambridge 1872)MKD01
         |--Typhochraestus Simon 1884S06
         |    |--T. barbatus (Koch 1880)S06
         |    `--T. spetsbergensis (Thorell 1871) [incl. Erigone oxycephala Koch 1880, E. provocans Cambr. 1877]S06
         |--Trachygnatha Kulcz. 1894S06
         |    |--T. dentata (Wid. 1834)S06
         |    `--T. taczanowskii (Cambr. 1873)S06
         |--Minicia Thorell 1875S06
         |    |--M. gibbosa Sörens. 1898S06
         |    `--M. marginella (Wider 1834)K02
         |--Poeciloneta Kulcz. 1894S06
         |    |--P. globosa (Wider 1841)BH02
         |    `--P. variegata (Blackwall 1841)MKD02
         |--Dunedinia Millidge 1988PVD10
         |    |--*D. denticulata Millidge 1988PVD19
         |    |--D. decolor Millidge 1988PVD10
         |    `--D. pullata Millidge 1988PVD10
         |--Laperousea de Dalmas 1917PVD10
         |    |--*L. blattifera (Urquhart 1887)PVD10 [=Linyphia blattiferNS00]
         |    |--L. cupidinea (Simon 1908)W76
         |    `--L. quindecimpunctata (Urquhardt 1893)W76
         |--Stemonyphantes Menge 1866 [Stemonyphantinae]S06
         |    |--S. conspersus (Koch 1879)MKD01
         |    |--S. lineatus (Linnaeus 1758) [incl. S. bucculentus]S06
         |    `--S. sibiricus (Grube 1861)MKD01
         |--Lophomma Menge 1867S06
         |    |--L. herbigradum (Blackw. 1854)S06
         |    |--L. nivicola Strand 1901S06
         |    `--L. punctatumRKD02
         |--Maso Simon 1884S06
         |    |--M. gallicus Simon 1894BBM02
         |    |--M. marxii (Keys. 1886)S06
         |    `--M. sundevalli (Westring 1851)MKD01
         |--Oreonetides Strand 1901S06
         |    |--O. abnormis [=Tmeticus abnormis]V09
         |    |--O. helsdingeni Eskov 1984MKD01
         |    `--O. vaginatus (Thorell 1872) [incl. Erigone adipata Koch 1872, E. macrochoera Thorell 1875]S06
         |--Macrargus Dahl 1886S06
         |    |--M. carpenteri (Cambridge 1894)K02
         |    |--M. multesimus (P.-Cambridge 1875)MKD01
         |    `--M. rufus (Wider 1830)SL14 [=Tmeticus rufusV09]
         |--Sintula Simon 1884S06
         |    |--S. affinis Kulcz. 1898S06
         |    |--S. cornigerRKD02
         |    |--S. obliviaP-C06
         |    `--S. spinigerSS02
         |--Labulla Simon 1884S06
         |    |--L. altioculata Keys. 1886S06
         |    |--L. machadoi Hormiga & Schaff 2005F05
         |    `--L. thoracica (Wider 1834)S06
         |--Drapetisca Menge 1866PVD10
         |    |--D. alteranda Chamberlin 1909DH14
         |    |--D. australis Forster 1955PVD10
         |    |--D. bicruris (Tu & Li 2006)DH14
         |    `--D. socialis (Sundevall 1833)DH14
         |--Caviphantes Oi 1960 [incl. Lessertiella Dumintrescu & Miller 1962]C90
         |    |--C. dobrogicusC90
         |    |--C. pseudosaxetorumC90
         |    |--C. samensisC90
         |    `--C. saxetorumC90
         |--Cnephalocotes Simon 1884S06
         |    |--C. laesus (Koch 1880)S06
         |    |--C. obscurus (Blackwall 1834)MKD01
         |    |--C. ophthalmicus Strand 1901S06
         |    `--C. pygmaeus Sörensen 1898S06
         |--Gonatium Menge 1866S06
         |    |--G. convexum Kulcz. 1885S06
         |    |--G. inflatum Sörens. 1898S06
         |    |--G. isabellinum (Koch 1841)S06
         |    `--G. rubens (Blackwall 1833)MKD01 [=Neriene rubensV09]
         |--Tmeticus Menge 1866S06
         |    |--T. affinis (Blackw. 1855)S06
         |    |--T. granulosus (Koch 1880)S06
         |    |--T. levinseni Sörens. 1898S06
         |    `--T. robustus (Westr. 1851)S06
         |--Entelecara Simon 1884S06
         |    |--E. acuminataV09
         |    |--E. congenera (P.-Cambridge 1879)MKD01
         |    |--E. erythropus (Westring 1851)BH02
         |    |--E. sombra (Chamberlin & Ivie 1947)MKD01
         |    `--E. trifrons (Cambr. 1863)S06
         |--Bolyphantes Koch 1837S06
         |    |--B. affinitatus Strand 1901S06
         |    |--B. alticeps (Sundevall 1833)MF11
         |    |--B. indexBV00
         |    |--B. luteolus (Blackwall 1833)K02
         |    `--B. sibiricus (Grube 1862)S06
         |--Diploplecta Millidge 1988PVD10
         |    |--*D. communis Millidge 1988PVD10
         |    |--D. adjacens Millidge 1988PVD10
         |    |--D. duplex Millidge 1988PVD10
         |    |--D. nuda Millidge 1988PVD10
         |    |--D. opaca Millidge 1988PVD10
         |    |--D. proxima Millidge 1988PVD10
         |    `--D. simplex Millidge 1988PVD10
         |--Maorineta Millidge 1988PVD10
         |    |--*M. tibialis Millidge 1988PVD10
         |    |--M. acerba Millidge 1988PVD10
         |    |--M. gentilis Millidge 1988PVD10
         |    |--M. minor Millidge 1988PVD10
         |    |--M. mollis Millidge 1988PVD10
         |    |--M. sulcata Millidge 1988PVD10
         |    `--M. tumida Millidge 1988PVD10
         `--Hilaira Simon 1884S06
              |--H. frigida (Thorell 1872) [incl. Erigone cryophila]S06
              |--H. gibbosa Tanasevitch 1982MKD01
              |--H. glacialis (Thorell 1871)S06
              |--H. herniosa (Thorell 1875)S06
              |--H. jamalensis Eskov 1981MKD01
              |--H. nubigena Hull 1911K02
              `--H. uncata (Cambr. 1872)S06
Linyphiidae incertae sedis:
  PityohyphantesMKD01
    |--P. costatus (Hentz 1850)WA02
    |--P. subarticus Chamberlin & Ivie 1943PS08
    `--P. phrygianus (Koch 1836)MKD01
  Drepanotylus uncatus (Cambridge 1873)K02
  Tallusia experta (Cambridge 1871)K02
  Taranucnus setosus (O.P.-Cambridge 1863)K02
  Monocephalus fuscipes (Blackwall 1836)BH02
  Panamomops sulcifrons (Wider 1834)BH02
  Syedra gracilis (Menge 1869)BH02
  Floronia bucculenta (Clerck 1757)BBM02
  Parapelecopsis nemoralis (Cambridge 1884)BBM02
  StyloctetorMKD01
    |--S. lehtineni Marusik & Tanasevitch 1999MKD01
    `--S. romanus (Cambridge 1872)BBM02
  Tapinopa longidens (Wider 1834)BBM02
  Saaristoa abnormis (Blackwall 1841)V09, BBM02
  GnathonariumMKD01
    |--G. dentatum (Wider 1834)MKD01 [=Gongylidium dentatumV09]
    `--G. suppositum (Kulczyński 1885)MKD01
  TroxochrusV09
    |--T. nasutusK08
    `--T. scabriculus (Westring 1851)J02 (see below for synonymy)
  Metopobactrus prominulus (Cambridge 1872)BBM02
  KaestneriaMKD01
    |--K. dorsalis (Wider 1834)MKD01 [=Bathyphantes dorsalisV09]
    `--K. pullata (P.-Cambridge 1863)MKD01
  Prinerigone vagans (Audouin 1826)MF11
  Thypochrestus digitatus (Cambridge 1872)BBM02
  Trichopterna cito (Cambridge 1872)BBM02
  Lepthothrix hardyi (Blackwall 1850)BBM02
  Trichoncoides piscator (Simon 1884)PO08
  TrichoncusMKD01
    |--T. hackmaniSS02
    |--T. hyperboreus Eskov 1992MKD01
    `--T. vasconicus Denis 1944MKD01
  NerieneMKD01
    |--N. clathrata (Sundevall 1830)BBM02 [=Linyphia clathrataV09]
    |--N. hammeni (van Helsdingen 1963)MKD01
    |--N. japonica (Oi 1960) [=Neolinyphia japonica]MKD01
    `--N. radiata (Walckenaer 1841)MKD01
  CentromeritaV09
    |--C. bicolor (Blackwall 1833)BBM02 [=Tmeticus bicolorV09]
    `--C. concinna (Thorell 1875)K02
  GongylidiellumPO08
    |--G. murcidumRKD02
    `--G. vivum (Cambridge 1875)BBM02
  MaroMKD01
    |--M. flavescens (P.-Cambridge 1873)MKD01
    |--M. lepidus Casemir 1961K02
    |--M. minutus Cambridge 1906K02
    `--M. sublestus Falconer 1915K02
  PelecopsisMKD01
    |--P. baicalensis Marusik, Koponen & Danilov 2001MKD01
    |--P. inedita (Pickard-Cambridge 1875)PO08
    |--P. janus Jocqué 1984J02
    |--P. mengei (Simon 1884)K02
    `--P. parallella (Wider 1834)MKD01 [=Lophocarenum parallelumV09]
  MicrargusSL14
    |--M. apertusRKD02
    |--M. herbigradus (Blackwall 1854)SL14
    `--M. subaequalis (Westring 1851)BH02
  Troglohyphantes strandiPN02
  Prolinyphia marginataJVHN90
  Aulacocyba subitaneaF75
  Sphecozone longipes (Strand 1908) [=Hypselistoides longipes]J98
  Bathylinyphia Eskov 1992MKD01
    `--*B. maior (Kulczyński 1885) (see below for synonymy)MKD01
  LasiargusMKD01
    |--L. hirsutus (Menge 1869)MKD01
    |--L. pilipes (Kulczyński 1908) [=Metopobactrus pilipes; incl. L. laricetorum Eskov 1989]MKD01
    `--L. zhui Eskov & Marusik 1994MKD01
  Silometopoides Eskov 1990MKD01
    |--*S. pampia (Chamberlin 1943) [=Minyriolus pampia]MKD01
    |--S. asiaticus (Eskov in Eskov & Marusik 1995) [=Silometopus asiaticus]MKD01
    |--S. koponeni (Eskov & Marusik 1994) [=Silometopus koponeni]MKD01
    |--S. mongolensis Eskov & Marusik 1992MKD01
    |--S. sibiricus (Eskov 1989) [=Silometopus sibiricus]MKD01
    |--S. sphagnicola Eskov & Marusik 1992MKD01
    |--S. tibialis (Heimer 1987) [=Metopobactrus tibialis]MKD01
    `--S. yodoensis (Oi 1960) [=Lophomma yodoensis]MKD01
  SilometopusMKD01
    |--S. ambiguus (Pickard-Cambridge 1905)PG08
    `--S. sachalinensis (Eskov & Marusik 1994) [=Silometopoides sachalinensis]MKD01
  Sauron Eskov in Eskov & Marusik 1995 [incl. Bothriopterna Miller & Zitnanska 1976 (n. n.)]MKD01
    |--*S. fissocornis Eskov 1995MKD01
    `--S. rayi (Simon 1881) [=Metopobactrus rayi; incl. Trichopterna fatrensis Miller 1966]MKD01
  Asiceratinops kolymensis Eskov 1992MKD01
  Asthenargoides kurenstchikovi Eskov 1993MKD01
  PacifiphantesMKD01
  Perlongipalpus pinipumilis Eskov & Marusik 1991MKD01
  Procerocymbium sibiricum Eskov 1989 [=P. sibiricus]MKD01
  Abacoproeces saltuum (Koch 1872)MKD01
  Abiskoa abiskoensis (Holm 1945)MKD01
  Concavocephalus rubens Eskov 1989MKD01
  Dactylopisthes video (Chamberlin & Ivie 1947)MKD01
  Decipiphantes decipiens (Koch 1879)MKD01
  Epibellowia pacifica (Eskov & Marusik 1992)MKD01
  Erigonidium graminicola (Sundevall 1830)MKD01
  Erigonoplus sibiricus Eskov & Marusik 1997MKD01
  Estrandia grandaeva (Keyserling 1886)MKD01
  Hylyphantes nigritus (Simon 1881)MKD01
  Hypomma bituberculatum (Wider 1834)MKD01 [=Dicyphus bituberculatusV09]
  IncestophantesMKD01
    |--I. calcaratus (Emerton 1909)PS08
    |--I. incestus (Koch 1879)MKD01
    `--I. kochiellus (Strand 1900)MKD01
  Ivielum sibiricum Eskov 1988MKD01
  Nenilinium asiaticum Eskov 1988MKD01
  Obscuriphantes pseudoobscurus (Marusik, Hippa & Koponen 1996)MKD01
  ParawubanoidesMKD01
    |--P. marusiki (Tanasevitch 1987)MKD01
    `--P. unicornis (P.-Cambridge 1873)MKD01
  PraestigiaMKD01
  Thaleria sukatchevae Eskov & Marusik 1992MKD01
  Tibioplus diversus (Koch 1897)MKD01
  Typhochrestoides baikalensis Eskov 1990MKD01
  Ummeliata sibirica (Eskov 1980)MKD01
  Walckenaerianus aimakensis Wunderlich 1995MKD01
  Wubanoides uralensis (Pakhorukov 1981)MKD01
    |--W. u. uralensisRL11
    `--W. u. lithodytes Schikora 2004RL11
  Yakutopus xerophilus Eskov 1990MKD01
  Zornella cultrigera (Koch 1879)MKD01
  DubiaraneaP-GGJ08
  ThaumatancusPO08
  Alioranus pastoralis (Pickard-Cambridge 1872)PO08
  Mecopisthes monticola Bosmans 1993PO08
  Australolinyphia Wunderlich 1976W76
    `--*A. remota Wunderlich 1976W76
  Knischatiria Wunderlich 1976W76
    `--*K. abnormis Wunderlich 1976W76
  Anacotyle setoensis Oi 1960W76
  Helophora insignis (Blackwall 1841)MF11
  Improphantes improbulus (Simon 1929)RL11
  AllomengeaD90
  WubanaD90
  Hylophantes graminicola [=Gongylidium graminicolum]V09
  Leptoroptrum robustumV09
  Baryphyma duffeyiJD-S07
  Proelauna humicolaJD-S07
  CallitrichiaJD-S07
  FrontinellinaJD-S07
  FrontinellaE95
    |--F. communisSG14
    `--F. pyramitelaE95
  ImpulsorS93
    |--I. mutilus Petrunkevitch 1958S93
    `--I. neglectus Petrunkevitch 1942S93
  Liticen setosus Petrunkevitch 1942S93
  MystagogusS93
    |--M. dubius Petrunkevitch 1942S93
    `--M. glaber Petrunkevitch 1942S93
  Custodela cheiracantha (Koch & Berendt 1854)S93
  Malleator niger Petrunkevitch 1942S93
  Meditrina circumvallata Petrunkevitch 1942S93
  Obnisus tenuipes Petrunkevitch 1942S93
  AntopiaP92
  ViocurusP92
  Malepellis extincta Petrunkevitch 1971P92
  PalaeolinyphiaP92

*Bathylinyphia maior (Kulczyński 1885) [=Bathyphantes maior, Bathylinyphia major (l. c.), Neriene major (l. c.); incl. Bathyphantes japonicus Oi 1980]MKD01

Ostearius melanopygius (Pickard-Cambridge 1879)W76 [incl. Erigone atriventer Urquhart 1887NS00, Neriene analis Simon 1894W76]

Troxochrus scabriculus (Westring 1851)J02 [=T. scabrosa (l. c.)BBM02; incl. T. cirrifrons (Cambridge 1871)BBM02, T. scabriculus cirrifronsJ02]

*Type species of generic name indicated

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