Lomechusini

Drusilla canaliculata, copyright Udo Schmidt.

Belongs within: Aleocharinae.
Contains: Zyras.

The Lomechusini are a diverse but poorly defined group of staphylinid beetles that have been recognised in the past (under the name Myrmedoniini) as including almost all Aleocharinae with 4,5,5-segmented tarsi. Many species possess mouthparts with extensive maxillary sinuses that extend caudally beyond the cardo, and an elongated galea and lacinia (Seevers 1957). Many, but not all, species are associated with ants and members of the subtribe Lomechusina possess prominent tufts of golden-yellow trichomes on the abdomen (Newton et al. 2001).

Characters (from Newton et al. 2001): Tarsi 4-5-5; metasternal process distinctly longer than very short mesosternal process; galea and lacinia of maxilla usually moderately to greatly elongated, length of galea equal to or greater than distance to base of galea from cardo; mesocoxae moderately to very broadly separated by broad meso- and metasternal processes.

<==Lomechusini [Myrmedoniini, Zyrasini]NT01
|--+--OrphnebiusLC20
| | |--O. dracoLC20
| | `--O. termitis Motschulsky 1859 [=Hygroptera termitis]S57
| `--+--Pella caliginosaLC20
| `--LomechusinaNT01
| |--LomechusaLC20
| | |--L. emarginataLC20
| | |--L. paradoxa [=Tachyporus (Lomechusa) paradoxus]G20
| | |--L. pubicollisLC20
| | `--L. strumosaA71
| `--Xenodusa Wasmann 1894 [incl. Pseudolomechusa Mann 1914]NT01
| |--X. cavaB14
| `--X. reflexa (Walker 1866)B14
`--Myrmedoniina [Myrmedoniae, Zyrasina]NT01
|--AtemelesS57
|--ZyrasNT01
|--Termozyras Cameron 1930S57
| `--*T. politus Cameron 1930S57
|--Urodonia Silvestri 1946S57
| `--*U. notabilis Silvestri 1946S57
|--Apalonia Casey 1906NT01
| `--A. seticornisNT01
|--Tetradonia Wasmann 1894 [incl. Chlorotusa Casey 1906]NT01
| `--T. megalops (Casey 1906)NT01
|--Trachyota Casey 1906NT01
| `--T. lativentrisNT01
`--Drusilla Leach 1819 [incl. Astilbus Dillwyn 1829, Myrmedonia Erichson 1837]NT01
|--*D. canaliculata (Fabricius 1787)S57, NT01 [=Astilbus canaliculatusC01]
|--D. cavicollis Casey 1906NT01
|--‘Myrmedonia’ clavigera Fauvel 1878O86
|--‘Myrmedonia’ humeralisC01
|--D. impressicollis [incl. D. sculpticollis]S57
|--D. incolaS57
|--‘Myrmedonia’ insignicornis Fauvel 1878O86
|--D. planaticollisS57
`--‘Myrmedonia’ tristis Lucas 1846E12

Lomechusini incertae sedis:
Dinocoryna Casey 1893 [incl. Ecitonusa Wasmann 1897]NT01
`--D. carolinensisNT01
Ecitocala Frank 1981NT01
`--E. rugosa Frank 1981NT01
Ecitonidia Wasmann 1900NT01
`--E. wheeleri Wasmann 1900NT01
Ecitoxenidia Wasmann 1909NT01
`--E. brevicornisNT01
Microdonia Casey 1893NT01
`--M. nitidiventrisNT01
Xesturida Casey 1906NT01
`--X. laevis Casey 1906NT01
SchistogeniaeS57
Termitonda Seevers 1957 [Termitondina]S57
|--*T. liberiae Seevers 1957S57
`--T. tachyporoides Seevers 1957S57
FeldinaS57
|--Felda Blackwelder 1952 [=Asticta Wasmann 1916 (preoc.)]S57
| `--*F. butteli (Wasmann 1916) [=*Asticta butteli]S57
|--Zunia Blackwelder 1952 [=Disticta Wasmann 1916 (preoc.)]S57
| `--*Z. capritermitis (Wasmann 1916) [=*Disticta capritermitis]S57
`--Termitobiella Wasmann 1916S57
`--*T. setipes Wasmann 1916S57
TermitozyrinaS57
|--Termitophagus Silvestri 1945S57
| `--*T. synterminus Silvestri 1945S57
|--Termitonusa Borgmeier 1950S57
| `--*T. sequax Borgmeier 1950S57
|--Iheringocantharus Bernhauer 1912S57
| `--*I. ypiranganus Bernhauer 1912S57
|--Termitosymbia Seevers 1957S57
| `--*T. nitida Seevers 1957S57
`--Termitozyras Seevers 1957S57
|--*T. adamsoni Seevers 1957S57
|--T. boliviae Seevers 1957S57
`--T. guianae Seevers 1957S57

*Type species of generic name indicated

References

[A71] Askew, R. R. 1971. Parasitic Insects. Heinemann Educational Books: London.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[LC20] Lü, L., C.-Y. Cai, X. Zhang, A. F. Newton, M. K. Thayer & H.-Z. Zhou. 2020. Linking evolutionary mode to palaeoclimate change reveals rapid radiations of staphylinoid beetles in low-energy conditions. Current Zoology 66 (4): 435–444.

[NT01] Newton, A. F., M. K. Thayer, J. S. Ashe & D. S. Chandler. 2001. Staphylinidae Latreille, 1802. In: Arnett, R. H., Jr & M. C. Thomas (eds) American Beetles vol. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia pp. 272–418. CRC Press: Boca Raton.

[O86] Olliff, A. S. 1886. A revision of the Staphylinidae of Australia. Part I. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 403–473, pl. 7.

[S57] Seevers, C. H. 1957. A monograph on the termitophilous Staphylinidae (Coleoptera). Fieldiana Zoology 40: 1–334.

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