Lytoceratida

Lytoceras fimbriatum, copyright Daderot.

Belongs within: Neoammonoidea.
Contains: Tetragonitidae, Gaudryceratidae, Crioceratites.

The Lytoceratida are a lineage of ammonites known from the Jurassic and Cretaceous. Their sutures are characterised by denticulated saddles with bipolar frilling (Wiedmann & Kullmann 1981).

Characters (from Engeser & Keupp 2002): Conch thin-walled, usually weakly sculptured. Suture line with primary sutural formula ELU2U1I; suture elements microphyll (strongly slit). Siphuncle usually reaching marginal position after one whorl; septal necks becoming prochoanitic at very early ontogenetic stage.

Lytoceratida [Lytoceratina]
| i. s.: Trachyphyllites costatusT81
|--Tetragonitoidea [Tetragonitaceae]EK02
| |--TetragonitidaeEK02
| `--GaudryceratidaeEK02
|--Crioceratitidae [Criocerataceae, Crioceratidae, Crioceratitinae, Himantoceratidae, Pedioceratidae, Toxoceratidae]EK02
| |--CrioceratitesEK02
| |--Eocrioceratites Wiedmann 1973WCH96
| | `--*E. rebillyi (Collignon 1962) [=Protancyloceras rebillyi]WCH96
| |--Megacrioceras Delanoy, Autran & Thomel 1987WCH96
| | `--*M. doublieri (Jaubert 1854) [=Ancyloceras doublieri]WCH96
| |--Hemicrioceras Spath 1924WCH96
| | `--*H. rude (Koenen 1902) [=Crioceras rude]WCH96
| |--Parancyloceras Spath 1924WCH96
| | `--*P. bidentatum (Koenen 1902) [=Crioceras bidentatum]WCH96
| |--Spinocrioceras Kemper 1973WCH96
| | `--*S. polyspinosum Kemper 1973WCH96
| |--Shasticrioceras Anderson 1938WCH96
| | `--*S. poniente Anderson 1938WCH96
| |--Simancyloceras Kemper 1973WCH96
| | |--*S. stolleyi Kemper 1973WCH96
| | `--S. pungueR81
| |--Pedioceras Gerhardt 1897 [incl. Pseudocrioceras Spath 1924]WCH96
| | |--*P. cundinamarcae Gerhardt 1897WCH96
| | `--‘Scaphites’ abichii Simonovich & Batsevich 1873 [=*Pseudocrioceras abichii]WCH96
| |--Menuthiocrioceras Collignon 1949WCH96
| | |--*M. lenoblei (Collignon 1949) [=Crioceras (*Menuthiocrioceras) lenoblei]WCH96
| | `--M. hourcquiWCH96
| |--Acantholytoceras Spath 1923 [incl. Pseudocrioceratites Egoian 1969]WCH96
| | |--*A. longispinum (Uhlig 1883) [=Hamites (Pictetia) longispinus]WCH96
| | `--*Pseudocrioceratites’ pseudoelegans Egoian 1969WCH96
| `--Aegocrioceras Spath 1924EK02 [incl. Bejucoceras Cantu-Chapa 1976WCH96]
| |--*A. capricornu (Roemer 1841) [=Hamites capricornu]WCH96
| |--A. bicarinatumWCH96
| |--A. compressum Rawson 1975EK02
| `--*Bejucoceras’ simplecostatum Cantu-Chapa 1976WCH96
`--LytocerataceaeP93
|--PleuroacanthitidaeP93
| |--Analytoceras articulatusP93
| `--Pleuroacanthites biformisP93
|--Derolytoceras [Derolytoceratidae]P93
| |--D. haueriP93
| `--D. tortumP93
|--NannolytoceratidaeP93
| |--AudaxlytocerasP93
| `--NannolytocerasP93
|--EctocentritidaeP93
| |--HolcolytocerasP93
| `--Ectocentrites Canavari 1888C81
| |--E. italicusC81
| `--E. petersiP93
`--Lytoceratidae [Ammonoceratidae, Carinolytoceratinae, Hemilytoceratinae, Lytoceratinae, Thysanoidae]EK02
|--Pterolytoceras Spath 1927WCH96
| `--*P. exoticum (Oppel 1863) [=Ammonites exoticus]WCH96
|--Eulytoceras Spath 1927WCH96
| `--*E. inaequalicostatum (Orbigny 1840) [=Ammonites inaequalicostatus]WCH96
|--Metalytoceras Spath 1927WCH96
| `--*M. triboleti (Uhlig 1902) [=Lytoceras triboleti]WCH96
|--Argonauticeras Anderson 1938WCH96
| `--*A. argonautarum (Anderson 1902) [=Lytoceras argonautarum]WCH96
|--Pictetia Uhlig 1883WCH96
| `--*P. astieriana (Orbigny 1842) [=Crioceras astierianum]WCH96
|--Carinolytoceras Wiedmann 1962WCH96
| `--*C. carinatum Wiedmann 1962WCH96
|--Hemilytoceras Spath 1927WCH96
| |--*H. immanis (Oppel 1865) [=Ammonites immanis]WCH96
| `--H. fraasiF12
|--Ammonoceratites Bowditch 1822 [incl. Ammonoceras Lamarck 1822]WCH96
| |--*A. lamarcki (Bowditch 1822) [=Ammonites lamarcki]WCH96
| |--A. ezoenseP93
| `--A. mahadeva (Stoliczka 1865)KK08
`--Lytoceras Suess 1865 (see below for synonymy)WCH96
|--*L. fimbriatum (Sowerby 1817) [=Ammonites fimbriatus]WCH96
|--L. cornucopiae (Young & Bird 1822)EK02
|--L. interlineatumP93
|--L. juilletiWK81
|--L. jurensisCDH04 [=Pachylytoceras jurenseC81; incl. P. aalenianum Buckman 1905C81]
|--‘Ammonites’ lineatus [incl. A. lineatus var. opalina]CDH04
|--‘Lobolytoceras’ siemensiL81
`--‘Pachylytoceras’ torulosum (Schübler in Zieten 1830)C81

Lytoceras Suess 1865 [incl. Biasaloceras Drushchits 1953, Cosmolytoceras Spath 1924, Crenilytoceras Buckman 1926, Dinolytoceras Beznosov 1956, Fimbrilytoceras Buckman 1918, Kallilytoceras Buckman 1921, Lobolytoceras Buckman 1923, Ophiceras Suess 1865, Orcholytoceras Buckman 1926, Saturnoceras Anderson 1938, Thysanoceras Hyatt 1867, Thysanolytoceras Buckman 1905, Trachylytoceras Buckman 1913, Valentolytoceras Beznosov 1956]WCH96

*Type species of generic name indicated

References

[C81] Callomon, J. H. 1981. Dimorphism in ammonoids. In: House, M. R., & J. R. Senior (eds) The Ammonoidea: The evolution, classification, mode of life and geological usefulness of a major fossil group pp. 257–273. Academic Press.

[CDH04] Callomon, J. H., D. T. Donovan & M. K. Howarth. 2004. F. A. Quenstedt’s trinomial nomenclature of Jurassic ammonites. Palaeontology 47 (4): 1063–1073.

[EK02] Engeser, T., & H. Keupp. 2002. Phylogeny of the aptychi-possessing Neoammonoidea (Aptychophora nov., Cephalopoda). Lethaia 35: 79–96.

[F12] Frýda, J. 2012. Phylogeny of Palaeozoic gastropods inferred from their ontogeny. In: Talent, J. A. (ed.) Earth and Life: Global biodiversity, extinction intervals and biogeographic perturbations through time pp. 395–435. Springer.

[KK08] Kennedy, W. J., & H. C. Klinger. 2008. Cretaceous faunas from Zululand and Natal, South Africa. The ammonite subfamily Lyelliceratinae Spath, 1921. African Natural History 4: 57–111.

[L81] Lehmann, U. 1981. Ammonites jaw apparatus and soft parts. In: House, M. R., & J. R. Senior (eds) The Ammonoidea: The evolution, classification, mode of life and geological usefulness of a major fossil group pp. 275–287. Academic Press.

[P93] Page, K. N. 1993. Mollusca: Cephalopoda (Ammonoidea: Phylloceratina, Lytoceratina, Ammonitina and Ancyloceratina). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 213–227. Chapman & Hall: London.

[R81] Rawson, P. F. 1981. Early Cretaceous ammonite biostratigraphy and biogeography. In: House, M. R., & J. R. Senior (eds) The Ammonoidea: The evolution, classification, mode of life and geological usefulness of a major fossil group pp. 499–529. Academic Press.

[T81] Tozer, E. T. 1981. Triassic Ammonoidea: classification, evolution and relationship with Permian and Jurassic forms. In: House, M. R., & J. R. Senior (eds) The Ammonoidea: The evolution, classification, mode of life and geological usefulness of a major fossil group pp. 66–100. Academic Press.

[WK81] Wiedmann, J., & J. Kullmann. 1981. Ammonoid sutures in ontogeny and phylogeny. In: House, M. R., & J. R. Senior (eds) The Ammonoidea: The evolution, classification, mode of life and geological usefulness of a major fossil group pp. 215–255. Academic Press.

[WCH96] Wright, C. W., J. H. Calloman & M. K. Howarth. 1996. Treatise on Invertebrate Paleontology pt L. Mollusca 4, revised vol. 4. Cretaceous Ammonoidea. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

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