Marshallena philippinarum, copyright Maurice Jay.

Belongs within: Horaiclavidae.

Marshallena is a genus of moderate- to large-sized conoids known from deep waters of the Indo-west Pacific (Powell 1966).

Characters (from Powell 1966): Shell of moderate to large size, 14–78 mm, fusiform-biconic, with medially angulate to subangulate turreted whorls, sculptured with closely spaced regular, narrowly crested axial folds, crossed by dense spiral cords and threads. Body-whorl slowly tapered to moderately long, slightly flexed, unnotched anterior canal. Sinus weak, a broad shallow concavity, occupying most of shoulder slope. Protoconch regular conic of three to five almost smooth whorls. Operculum small, thin, corneous, ovated, with subterminal nucleus that curves inward toward columella. Radula a pair of simple wishbone-type marginals; basal limbs bridged by thin plate.

<==Marshallena Allan 1927BK11 [incl. Sugitania Kuroda 1958 non Matsumura 1926P66, Sugitanitoma Kuroda 1959P66]
    |--*M. neozelanica (Suter 1917) [=Daphnella neozelanica; incl. Belophos incertus Marshall 1919]P66
    |--M. anomala Powell 1942P66
    |--M. austrotomoides Powell 1931P66
    |--M. carinaria Powell 1935P66
    |--M. celsa Marwick 1931P66
    |--M. curtata (Marwick 1926) [=Turricula curtata]P66
    |--M. decens Marwick 1931P66
    |--M. esdailei (Marwick 1926) [=Turricula esdailei]P66
    |--M. gracilispira Powell 1969MG-H11
    |--M. impar Powell 1942P66
    |--M. ‘lepta’ (Watson 1881)P66 [=Pleurotoma (Surcula) lepta non P. lepta Edwards 1861P66, Turricula leptaH22]
    |--M. nereis (Smith 1906) [=Pleurotoma nereis]P66
    |    |--M. n. nereisP66
    |    `--M. n. ‘reticulata’ (Kuroda 1959) non Sugitania reticulata Kuroda 1958P66
    |--M. nierstraszi (Schepman 1913) [=Surcula nierstraszi]P66
    |--M. ovata (Marshall 1917) [=Daphnella ovata]F27
    |--M. philippinarum (Watson 1882)BK11
    |--M. reticulata (Kuroda 1958) [=*Sugitania reticulata, *Sugitanoma reticulata]P66
    `--M. serotina (Suter 1917) [=Surcula serotina]P66

*Type species of generic name indicated


[BK11] Bouchet, P., Y. I. Kantor, A. Sysoev & N. Puillandre. 2011. A new operational classification of the Conoidea (Gastropoda). Journal of Molluscan Studies 77: 273–308.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[H22] Hedley, C. 1922. A revision of the Australian Turridae. Records of the Australian Museum 13 (6): 213–359, pls 42–56.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[P66] Powell, A. W. B. 1966. The molluscan families Speightiidae and Turridae: an evaluation of the valid taxa, both Recent and fossil, with lists of characteristic species. Bulletin of the Auckland Institute and Museum 5: 1–184, pls 1–23.

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