Melyridae

Melyris bicolor, copyright Efim.

Belongs within: Cleroidea.
Contains: Listrini, Dasytes, Malachiinae.

The Melyridae are a group of relatively soft-bodied beetles that are usually predacious or scavengers as larvae but commonly flower-feeders as adults. The elytra are commonly reduced in the subfamily Malachiinae but not in the subfamilies Melyrinae and Dasytinae. Melyrinae are often brightly coloured (bicoloured as in Malachiinae or metallic blue or green) whereas Dasytinae are usually uniformly black, brown or yellow. Dasytinae have the first tarsomere on all legs as long as or longer than the second whereas it is shorter in Melyrinae. Within the Dasytinae, members of the tribe Dasytini typically have tarsal claws with a basal tooth, and tarsomere 4 shorter than 3 on all legs (Mayor 2002).

Characters (from Lawrence & Britton 1991): Oblong to elongate, somewhat flattened, soft-bodied, usually clothed with decumbent hairs and scattered erect setae, either uniformly black or grey or bicoloured with red or yellow and blue or black . Labrum well sclerotised; tarsal claws toothed or provided with a pair of fleshy appendages; elytra sometimes very short, exposing several abdominal tergites; eversible vesicles often on prothorax, mesothorax and abdomen. Larvae elongate, slightly flattened, lightly sclerotised, except for head, T9 and thoracic plates, red or pink in colour; head with long epicranial stem, no median endocarina, and lightly sclerotised gular and para-gular areas, so that head capsule appears to be open behind ventral mouthparts; mesal appendage at base of mandible.

<==Melyridae [Attalomimidae, Carphuridae, Gietellidae, Malachiidae]
|--MelyrinaeMW15
| |--CerralusM02
| |--AnthodromiusM02
| |--ChalchasM02
| |--ArthrobrachusM02
| |--AstylomorphusM02
| |--MicrozygiaM02
| |--FalsomelyrisM02
| |--PseudozygiaM02
| |--ProcerallusM02
| |--Astylus [Astylini]MW15
| | `--A. atromaculatus (Blanchard 1843)B14
| |--Melyrodes Gorham 1882 [incl. Alymeris Casey 1895]M02
| | |--M. basalis (LeConte 1852) [=Dasytes basalis]B14
| | |--M. cribratus (LeConte 1852)M02
| | `--M. floridiana (Casey 1895)M02
| `--Melyris Fabricius 1775M02
| |--M. bicolorG89
| |--M. corrosaG01
| |--M. festivaG01
| |--M. rubripes Lucas 1846E12
| |--M. rufiventrisG01
| `--M. viridisMW15
`--+--Dasytinae [Danaceinae, Dasytidae]MW15
| | i. s.: Amecocerus Solier 1849BM76, M02
| | `--A. cervicalisBM76
| |--ListriniM02
| `--DasytiniM02
| |--DasytesB14
| |--Enicopus (Parahenicopus) lopezcoloni Bahillo de la Puebla 2005M02, F05
| |--DivalesM02
| |--PsilothrixM02
| |--AllotarsusM02
| |--DolichophronM02
| |--HauseriaM02
| |--AllotarsodytesM02
| |--TrochantodonM02
| |--GraellsinusM02
| |--Dasytellus Casey 1895M02
| |--Dasytastes Casey 1895M02
| |--Vectura Casey 1895 [incl. Pseudallonyx Casey 1895]M02
| | `--V. longicepsM02
| |--Leptovectura Casey 1895M02
| | `--L. adspersa Casey 1895M02
| |--Amphivectura Blaisdell 1938M02
| | `--A. monticola (Blaisdell 1934)M02
| |--Hoppingiana Blaisdell 1924M02
| | |--H. hudsonicus (LeConte 1866)M02
| | `--H. nitida Hatch 1962M02
| |--Dolichosoma Stephens 1830M02
| | |--D. foveicolle (Kirby 1837)M02
| | |--D. lineareC01
| | `--D. tenuiforme Horn 1880M02
| |--Vecturoides Fall 1930 [incl. Menovectura Blaisdell 1931]M02
| | |--V. albicans (Casey 1895) [=Vectura albicans]M02
| | |--V. pseudonycha Fall 1930M02
| | `--V. serrulata (Blaisdell 1931)M02
| |--Enallonyx Wolcott 1944M02
| | |--E. cinerescens (Fall 1930)M02
| | |--E. denudatus (Casey 1895)M02
| | |--E. disjunctus (Casey 1895)M02
| | `--E. sculptilis (LeConte 1859)M02
| `--Eschatocrepis LeConte 1862M02
| `--E. constrictus (LeConte 1852)M02
| |--E. c. constrictusM02
| |--E. c. desertus Blaisdell 1931M02
| |--E. c. nigripes Blaisdell 1921M02
| `--E. c. riversidensis Howell 1985M02
`--MalachiinaeMW15

Melyridae incertae sedis:
SimoderusMC13
ChalicorusG01
|--C. collarisG01
|--C. formicarius Gorham 1901G01
`--C. vinulusG01
MyrmecospectaG01
HapalochrusG01
|--H. aerosus Gorham 1901G01
|--H. cinctusM86
`--H. mashunus Gorham 1901G01
HedybiusG01
|--H. albipennis Gorham 1901G01
|--H. amoenusG00
|--H. anceps Gorham 1900G00
|--H. lividusG01
|--H. marshalli Gorham 1900G00
|--H. superciliosusG01
`--H. variicornisG01
DasytinaP92
MicrojulistusP92
Zygia Fabricius 1775P92, G76
`--Z. oblonga (Fabricius 1775)L02, M02 [=Melyris (Zyga) oblongaG20]
Dinometopus Gorham 1900G00
`--*D. natalensis Gorham 1900G00
CephaloncusG00
ApalochrusG00
|--A. azureusG00
`--A. nitens Gorham 1900G00
Gietella [Gietellinae]M02
`--G. fortunata Constantin & Menier 1987M02

*Type species of generic name indicated

References

[BM76] Bohart, R. M., & A. S. Menke. 1976. Sphecid Wasps of the World. University of California Press: Berkeley.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature’s gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[G89] Gestro, R. 1889. Viaggio ab Assab nel Mar Rosso dei signori G. Doria ed O. Beccari con il R. Avviso “Esploratore” dal 16 Novembre 1879 al 26 Febbraio 1880.—IV. Coleotteri. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a, 7: 5–72.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[G00] Gorham, H. S. 1900. Descriptions of new genera and species of Coleoptera from South and West Africa, of the section Serricornia, and of the families Erotylidae, Endomychidae, and Languriidae. Annals and Magazine of Natural History, series 7, 5: 71–94.

[G01] Gorham, H. S. 1901. Descriptions of genera and species of Coleoptera from South Africa (continued). Annals and Magazine of Natural History, series 7, 7: 349–365.

[G76] Grasshoff, M. 1976. Zur Taxonomie und Nomenklatur mitteleuropäischer Radnetzspinnen der Familie Araneidae (Arachnida: Araneae). Senckenbergiana Biologica 57 (1–3): 143–154.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[MC13] Majer, J. D., S. K. Callan, K. Edwards, N. R. Gunawardene & C. K. Taylor. 2013. Baseline survey of the terrestrial invertebrate fauna of Barrow Island. Records of the Western Australian Museum Supplement 83: 13–112.

[M02] Mayor, A. J. 2002. Melyridae Leach 1815. In: Arnett, R. H., Jr, M. C. Thomas, P. E. Skelley & J. H. Frank (eds) American Beetles vol. 2. Polyphaga: Scarabaeoidea through Curculionoidea pp. 281–304. CRC Press.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part IV. Proceedings of the Linnean Society of New South Wales, series 2, 1 (2): 259–380.

[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

[P92] Poinar, G. O., Jr. 1992. Life in Amber. Stanford University Press: Stanford.

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