Metacopina

Left valve, dorsal view and interior of left valve of Costatia posneri, from Benson et al. (1961).

Belongs within: Ostracoda.
Contains: Platycopina, Healdiidae, Thlipsuridae.

The Metacopina are a group of ostracods in which the carapace valves ancestrally bear muscle scars arranged as a circular aggregate of many scars (Benson et al. 1961).

Characters (from Benson et al. 1961): Hinge distinct, simple to tripartite; muscle scar consisting of secondary scars assembled in a compact group; inner lamella narrow, poorly developed or unknown.

<==Metacopina S61b
    |–+–Platycopina S61b
    |  `–Thlipsuroidea [Healdiacea, Thlipsuracea, Thlipsuridacea] S61b
    |       |  i. s.: Necrateria Gründel 1962 S82
    |       |           `–N. rectagona (Gründel 1962) [=Triplacera (Necrateria) rectagona] S82
    |       |–Healdiidae HM04
    |       |–Thlipsuridae S61b
    |       `–Krausellidae BB61
    |            |–Krausella Ulrich 1894 [incl. Basslerites Teichert 1937 non Howe 1937, Rayella Teichert 1939] BB61
    |            |    `–*K. inaequalis Ulrich 1894 BB61
    |            |–Cooperatia Tolmachoff 1937 [=Cooperia Tolmachoff 1926 non Ransom 1907] BB61
    |            |    `–*C. granum (Tolmachoff 1926) [=*Cooperia granum] BB61
    |            `–Janusella Roth 1929 BB61
    |                 `–*J. biceratina Roth 1929 BB61
    `–Quasillitacea S61b
         |  i. s.: Unicornites Pokorný 1950 BB61
         |           `–*U. homeomorphus Pokorný 1950 BB61
         |–Ropolonellidae BB61
         |    |–Varix [=Varis (l. c.)] M61b
         |    |–Ropolonellus Van Pelt 1933 BB61
         |    |    `–*R. papillatus Van Pelt 1933 BB61
         |    |–Rudderina Coryell & Malkin 1936 BB61
         |    |    `–*R. extensa Coryell & Malkin 1936 BB61
         |    |–Varicobairdia Pokorný 1950 BB61
         |    |    `–*V. kettneri (Pokorný 1950) [=Bairdia (*Varicobairdia) kettneri] BB61
         |    `–Plagionephrodes Morey 1935 (n. d.) M61b, BB61
         |         `–*P. uninodosus Morey 1935 BB61
         |–Bufinidae BB61
         |    |–Punctomosea Stover 1956 BB61
         |    |    `–*P. cristata (Swartz & Oriel 1948) [=Thrallella cristata] BB61
         |    |–Aurigerites Roundy 1926 S82
         |    |    |–*A. texanus Roundy 1926 BB61
         |    |    `–A. obernitzensis Gründel 1942 S82
         |    |–Bufina Coryell & Malkin 1936 [incl. Parabufina Smith 1956] BB61
         |    |    |–*B. elata Coryell & Malkin 1936 BB61
         |    |    `–B. bicornuta (Ulrich 1891) [=Moorea bicornuta] BB61
         |    `–Bythocyproidea Stewart & Hendrix 1945 BB61
         |         |–*B. sanduskyensis Stewart & Hendrix 1945 BB61
         |         `–B. eriensis BB61
         `–Quasillitidae [Graphiodactylidae] S61b
              |–Eriella Stewart & Hendrix 1945 BB61
              |    `–*E. robusta Stewart & Hendrix 1945 BB61
              |–Euglyphella Warthin 1934 BB61
              |    `–*E. sigmoidalis (Jones 1890) [=Strepula sigmoidalis] BB61
              |–Graphiadactylis Roth 1929 BB61 (see below for synonymy)
              |    `–*G. arkansana (Girty 1910) [=Kirkbya lindahli arkansana] BB61
              |–Jenningsina Coryell & Malkin 1936 BB61
              |    `–*J. catenulata (Van Pelt 1933) [=Graphiodactylus catenulatus] BB61
              |–Scalptina Stover 1956 BB61
              |    `–*S. incuda Stover 1956 BB61
              |–Paragraphylus Coryell & Rozansky 1942 (n. d.) M61b, BB61
              |    `–*P. reticulatus Coryell & Rozansky 1942 BB61
              |–Allostraca Ulrich & Bassler 1932 (n. d.) M61b, BB61
              |    `–*A. fimbriata Ulrich & Bassler 1932 BB61
              |–Quasillites Coryell & Malkin 1936 (see below for synonymy) BB61
              |    |–*Q. obliquus Coryell & Malkin 1936 BB61
              |    `–*Lucasella’ mundula Stewart 1936 (n. d.) BB61
              `–Costatia Polenova 1952 BB61
                   |–*C. posneri Polenova 1952 BB61
                   `–C. cavernosa BB61

Graphiadactylis Roth 1929 BB61 [=Graphiodactyllis (l. c.) M61b, Graphiodactylus Roth 1929 BB61; incl. Bassleria Harlton 1929 BB61]

Quasillites Coryell & Malkin 1936 [incl. Allostracites Přibyl 1953, Burlella Coryell & Booth 1933, Lucasella Stewart 1936, Paracythere Bassler 1932 non Müller 1894, Spinovina Coryell & Malkin 1936] BB61

*Type species of generic name indicated

REFERENCES

[BB61] Benson, R. H., J. M. Berdan, W. A. van den Bold, T. Hanai, I. Hessland, H. V. Howe, R. V. Kesling, S. A. Levinson, R. A. Reyment, R. C. Moore, H. W. Scott, R. H. Shaver, I. G. Sohn, L. E. Stover, F. M. Swain & P. C. Sylvester-Bradley. 1961. Systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q99–Q421. Geological Society of America and University of Kansas Press.

[HM04] Hoare, R. D., & G. K. Merrill. 2004. A Pennsylvanian (Morrowan) ostracode fauna from Texas. Journal of Paleontology 78 (1): 185–204.

[M61] Moore, R. C. 1961. Supplement—USSR treatise on Ostracoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q422–Q429. Geological Society of America and University of Kansas Press.

[S61] Scott, H. W. 1961. Classification of Ostracoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q74–Q92. Geological Society of America and University of Kansas Press.

[S82] Shi C.-G. 1982. Some early Carboniferous ostracodes from Nylam, Xizang (Tibet). Acta Palaeontologica Sinica 21 (3): 309–314.

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